The story of the universe. Volume 3 (of 4) : $b The earth's garment : flora
History
The story of the universe. Volume 3 (of 4) : $b The earth's garment : flora
Astronomy; Earth (Planet); Natural history
Petals in ordinary flowers are intended to secure the attention
of insects; to wind-fertilized blossoms, having no occasion for
visitors, they are unnecessary. So far from an advantage, the
presence of a corolla would exclude the wind from the essential
organs. Accordingly, petals are either absent altogether or reduced
to rudimentary proportions. The calyx is also much reduced, and
in some flowers is dispensed with entirely. Comparatively few
anemophilous flowers possess both sets of floral envelopes.
Plantago is, however, dichlamydeous, but its chaffy petals afford
incontrovertible evidence of degeneration from the entomophilous
condition.
The stigma in the wind-fertilized class is highly specialized, and
much larger relatively to the other parts of the flower than is
the case with entomophilous blossoms. It is commonly penicillate,
consisting of a tuft of hairs, as in nettle; feathery, as in grasses;
or elongated and thread-like, as in plantago and the rushes. The
spirally twisted stigmas of the last-mentioned flowers are beautiful
objects when examined with a pocket lens. The larger the surface
which the stigma presents to the wind, the greater are the chances
of pollination. Its fine fringes of papillose hairs are also well
calculated to entangle the pollen-grains, while the viscid secretion
serves to retain them when caught. This adaptation may be seen in the
common rye grass; each tiny blossom as it expands hangs out its two
white, feathery stigmas from the sides of the spikelet, reminding
one of a fisherman spreading out his nets, or a sailor his studding
sails to catch the favoring breeze. At the time of fertilization the
dock, too, thrusts out its three little brush-like stigmas between
the lobes of the perianth. It is instructive to compare these
wind-fertilized flowers of Rumex with those of the nearly allied
genus Polygonum, which is entomophilous. The perianth of the latter
is rose-colored; the stigmas are included within it, never exserted
as in the dock--they are not at all brush-like or feathery, but in
the form of little knobs; the stamens and flower-stalks are rigid;
moreover, the various species of Polygonum secrete nectar and are
frequented by many different insects. Stigmas are entirely absent in
the gymnospermous division, but in most Coniferæ the ovule at the
time of flowering secretes a drop of liquid, and the pollen-grains
caught on it are, as the fluid gradually evaporates, stranded on
the nucleus of the ovule. The ovule of the larch is provided with
elongated papillæ, functionally equivalent to a stigma.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account