The story of the universe. Volume 3 (of 4) : $b The earth's garment : flora
History
The story of the universe. Volume 3 (of 4) : $b The earth's garment : flora
Astronomy; Earth (Planet); Natural history
The internal anatomy of the Lycopodiaceæ is somewhat complex, but
their external organization is simple. A club moss consists of a
cylindrical stem covered with overlapping leaves, spirally arranged,
of small size relatively to the stem, and always simple or undivided.
The stem branches in a peculiar forked manner, which gives the
plant its characteristic candelabra-like form. Existing lycopods
are creeping plants, seldom exceeding two feet in height, but many
extinct species attained the dimensions of large trees. On the ends
of certain branches the leaves are crowded together, giving the
terminal portion of each shoot some resemblance to a pine-cone. The
crowded leaves on this portion bear, on their upper surfaces, little
sacs called sporangia. Certain of these sacs contain very numerous
small, rounded bodies, the microspores; others have fewer spores
of larger size, distinguished as macrospores. Sacs containing the
small male spores are termed microsporangia; those having the large
female spores, macrosporangia. When ripe, a sporangium bursts and
discharges its spores, which are scattered by the wind. Should a
spore alight on a favorable spot, it germinates after a time and
gives rise to a structure called a prothallus, which is really an
independent plant. This stage in the life-history of a cryptogam is,
however, much better seen in ferns, where the prothallus is entirely
expelled from the spore and attains a higher degree of independent
development. The prothallus throws out root-hairs, nourishes
itself and grows, but the leaf-like form it assumes bears not the
remotest resemblance to the parent fern from which it sprang. This
phenomenon, characteristic of the higher cryptogams, is known as the
“alternation of generations,” or “alternate generations.” Similar
phases are observed in certain animals, the medusæ or jelly fishes,
for example. In the course of its development, a fern passes through
two distinct phases; first, the spore-bearing stage or sporophyte,
represented by the fern frond; second, the egg-bearing stage, the
oöphyte or prothallus. As we ascend in the scale of vegetable life,
the egg-bearing or sexual generation diminishes in importance,
while the sporophyte preponderates more and more. In club mosses,
the prothallus has all but lost its independence; in the case of
the selaginella it is formed almost entirely within the spore, only
a small part being extruded when the spore ruptures. Some of the
lycopods are inosporous--that is, they have, like the ferns, but
one kind of spore. Where this is the case, the prothallus developed
from the spore bears two sets of sexual organs; the prothallus of
one of the heterosporous cryptogams, on the other hand, produces
sexual organs of one kind only. Antheridia appear on the prothallus
developed from a small spore; archegonia on that from a large one.
The former are the male organs, and from them are emitted numerous
antherozoids, minute ciliated bodies, which swarm over damp surfaces
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