The story of the universe. Volume 4 (of 4) : $b The earth's creatures : fauna
History
The story of the universe. Volume 4 (of 4) : $b The earth's creatures : fauna
Astronomy; Earth (Planet); Natural history
These considerations supply us with a key to many of those apparent
anomalies which can not fail to strike us in viewing the dispositions
of the parts of the skeleton and the remarkable inversion they appear
to have undergone, when compared with the usual arrangement. We
find, however, on a more attentive examination, that all the bones
composing the skeleton in other vertebrated animals exist also in
the tortoise; and that the bony case which envelops all the other
parts is really formed by an extension of the spinous processes of
the vertebræ and ribs on the one side and of the usual pieces which
compose the sternum on the other. The upper and lower plates thus
formed are united at their edges by expansions of the sternocostal
appendices, which become ossified. Thus, no new element has been
created; but advantage has been taken of those already existing in
the general type of the vertebrata, to modify their forms by giving
them different degrees of relative development, and converting them,
by these transformations, into a mechanism of a very different kind,
and subservient to other objects than those to which they are usually
applied. It is scarcely possible to have stronger proofs, if such
were wanting, of the unity of plan which has regulated the formation
of all animal structures than those afforded by the skeleton of the
tortoise.
The first step taken to secure the relative immobility of the trunk
is to unite in one rigid, bony column all its vertebræ, and to allow
of motion only in those of the neck and of the tail. The former,
accordingly, are all anchylosed together, leaving, indeed, traces
of their original forms as separate vertebræ, but exhibiting no
sutures at the place of junction. The canal for the spinal marrow is
preserved, as usual, above the bodies of these coalesced vertebræ,
and is formed by their united leaves; the arches being completed by
the spinous processes. But these processes do not terminate in a
crest as usual; they are further expanded in a lateral direction,
forming flat pieces along the back, which are united to one another
by sutures, and which are also joined to the expanded ribs, so as to
form the continuous plane surface of the carapace. The transverse
processes of the vertebræ are well marked, but, though firmly united
to the ribs, do not give rise to them; for the ribs, which are
flattened and expanded, so as to touch one another along their whole
length, are inserted below, between the bodies of every two adjoining
vertebræ; while above they are united by suture with the plates of
the spinous processes. This change in the situation of the ribs is
the consequence of the change in their office. When designed to be
very movable, we find them attached either to the extremities of
the transverse processes or to the articular surfaces of a single
vertebra; but where solidity and security are aimed at, they are
always inserted between the bodies of two vertebræ. It is remarkable,
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