light. 22 Average of above seven large lop-eared
rabbits 1268 132 11 per cent. too light.
In the upper half of Table 3 I have given the measurements of the skull
of ten wild rabbits; and in the lower half, of eleven thoroughly
domesticated kinds. As these rabbits differ so greatly in size, it is
necessary to have some standard by which to compare the capacities of
their skulls. I have selected the length of skull as the best standard,
for in the larger rabbits it has not, as already stated, increased in
length so much as the body; but as the skull, like every other part,
varies in length, neither it nor any other part affords a perfect
standard.
In the first column of figures the extreme length of the skull is given
in inches and decimals. I am aware that these measurements pretend to
greater accuracy than is possible; but I have found it the least
trouble to record the exact length which the compass gave. The second
and third columns give the length and weight of body, whenever these
observations were made. The fourth column gives the capacity of the
skull by the weight of small shot with which the skulls were filled;
but it is not pretended that these weights are accurate within a few
grains. In the fifth column the capacity is given which the skull ought
to have had by calculation, according to the length of skull, in
comparison with that of the wild rabbit No. 1; in the sixth column the
difference between the actual and calculated capacities, and in the
seventh the percentage of increase or decrease, are given. For
instance, as the wild rabbit No. 5 has a shorter and lighter body than
the wild rabbit No. 1, we might have expected that its skull would have
had less capacity; the actual capacity, as expressed by the weight of
shot, is 875 grains, which is 97 grains less than that of the first
rabbit. But comparing these two rabbits by the length of their skulls,
we see that in No. 1 the skull is 3·15 inches in length, and in No. 5
2·96 inches in length; according to this ratio, the brain of No. 5
ought to have had a capacity of 913 grains of shot, which is above the
actual capacity, but only by 38 grains. Or, to put the case in another
way (as in column vii), the brain of this small rabbit, No. 5, for
every 100 grains of weight is only 4 grains too light,—that is, it
ought, according to the standard rabbit No. 1, to have been 4 per cent
heavier. I have taken the rabbit No. 1 as the standard of comparison
because, of the skulls having a full average length, this has the least
capacity; so that it is the least favourable to the result which I wish
to show, namely, that the brain in all long-domesticated rabbits has
decreased in size, either actually, or relatively to the length of the
head and body, in comparison with the brain of the wild rabbit. Had I
taken the Irish rabbit, No. 3, as the standard, the following results
would have been somewhat more striking.
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