What, then, are we to conclude from this tendency in all the chief
domestic races, both when purely bred and more especially when
intercrossed, to produce offspring of a blue colour, with the same
characteristic marks, varying in the same manner, as in _Columbia
livia_? If we admit that these races are all descended from _C. livia,_
no breeder will doubt that the occasional appearance of blue birds thus
characterised is accounted for on the well-known principle of “throwing
back” or reversion. Why crossing should give so strong a tendency to
reversion, we do not with certainty know; but abundant evidence of this
fact will be given in the following chapters. It is probable that I
might have bred even for a century pure black Barbs, Spots, Nuns, white
Fantails, Trumpeters, etc., without obtaining a single blue or barred
bird; yet by crossing these breeds I reared in the first and second
generation, during the course of only three or four years, a
considerable number of young birds, more or less plainly coloured blue,
and with most of the characteristic marks. When black and white, or
black and red birds, are crossed, it would appear that a slight
tendency exists in both parents to produce blue offspring, and that
this, when combined, overpowers the separate tendency in either parent
to produce black, or white, or red offspring.
If we reject the belief that all the races of the pigeon are the
modified descendants of _C. livia,_ and suppose that they are descended
from several aboriginal stocks, then we must choose between the three
following assumptions: firstly, that at least eight or nine species
formerly existed which were aboriginally coloured in various ways, but
have since varied in exactly the same manner so as to assume the
colouring of _C. livia_; but this assumption throws not the least light
on the appearance of such colours and marks when the races are crossed.
Or secondly, we may assume that the aboriginal species were all
coloured blue, and had the wing-bars and other characteristic marks of
_C. livia,_—a supposition which is highly improbable, as besides this
one species no existing member of the Columbidæ presents these combined
characters; and it would not be possible to find any other instance of
several species identical in plumage, yet as different in important
points of structure as are Pouters, Fantails, Carriers, Tumblers, etc.
Or lastly, we may assume that all the races, whether descended from _C.
livia_ or from several aboriginal species, although they have been bred
with so much care and are so highly valued by fanciers, have all been
crossed within a dozen or score of generations with _C. livia,_ and
have thus acquired their tendency to produce blue birds with the
several characteristic marks. I have said that it must be assumed that
each race has been crossed with _C. livia_ within a dozen, or, at the
utmost, within a score of generations; for there is no reason to
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