Another case of analogous variation is the occurrence of spangled
sub-breeds of Hamburgh, Polish, Malay, and Bantam fowls. Spangled
feathers have a dark mark, properly crescent-shaped, on their tips;
whilst pencilled feathers have several transverse bars. The spangling
cannot be due to reversion to _G. bankiva_; nor does it often follow,
as I hear from Mr. Tegetmeier, from crossing distinct breeds; but it is
a case of analogous variation, for many gallinaceous birds have
spangled feathers,—for instance, the common pheasant. Hence spangled
breeds are often called “pheasant”-fowls. Another case of analogous
variation in several domestic breeds is inexplicable; it is, that the
chickens, whilst covered with down, of the black Spanish, black Game,
black Polish, and black Bantam, all have white throats and breasts, and
often have some white on their wings.[31] The editor of the ‘Poultry
Chronicle’[32] remarks that all the breeds which properly have red
ear-lappets occasionally produce birds with white ear-Tappets. This
remark more especially applies to the Game breed, which of all comes
nearest to the _G. bankiva_; and we have seen that with this species
living in a state of nature, the ear-lappets vary in colour, being red
in the Malayan countries, and generally, but not invariably, white in
India.
In concluding this part of my subject, I may repeat that there exists
one widely-ranging, varying, and common species of Gallus, namely, _G.
bankiva,_ which can be tamed, produces fertile offspring when crossed
with common fowls, and closely resembles in its whole structure,
plumage, and voice the Game breed; hence it may be safely ranked as the
parent of this, the most typical domesticated breed. We have seen that
there is much difficulty in believing that other, now unknown, species
have been the parents of the other domestic breeds. We know that all
the breeds are most closely allied, as shown by their similarity in
most points of structure and in habits, and by the analogous manner in
which they vary. We have also seen that several of the most distinct
breeds occasionally or habitually closely resemble in plumage _G.
bankiva,_ and that the crossed offspring of other breeds, which are not
thus coloured, show a stronger or weaker tendency to revert to this
same plumage. Some of the breeds, which appear the most distinct and
the least likely to have proceeded from _G. bankiva,_ such as Polish
fowls, with their protuberant and little ossified skulls, and Cochins,
with their imperfect tail and small wings, bear in these characters the
plain marks of their artificial origin. We know well that of late years
methodical selection has greatly improved and fixed many characters;
and we have every reason to believe that unconscious selection, carried
on for many generations, will have steadily augmented each new
peculiarity, and thus have given rise to new breeds. As soon as two or
three breeds were once formed, crossing would come into play in
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