Finally, I have not examined a sufficient number of skeletons to say
whether any of the foregoing differences, except in the skull, are
characteristic of the several breeds. Apparently some differences are more
common in certain breeds than in others,--as an additional rib to the
fourteenth cervical vertebra in Hamburghs and Games, and the breadth of the
end of the pubic bone in Cochins. Both skeletons of the Sultan fowl had
eight dorsal vertebræ, and the end of the scapula in both was somewhat
attenuated. In the skull, the deep medial furrow in the frontal bones and
the vertically elongated occipital foramen seem to be characteristic of
Cochins; as is the great breadth of the frontal bones in Dorkings; the
separation and open spaces between the tips of the ascending branches of
the premaxillaries and nasal bones, as well as the front part of the skull
being but little depressed, characterise Hamburghs; the globular shape of
the posterior part of the skull seems to be characteristic of laced
Bantams; and lastly, the protuberance of the skull with the ascending
branches of the premaxillaries partially aborted, together with the other
differences before specified, are eminently characteristic of Polish and
other Crested fowls.
But the most striking result of our examination of the skeleton is the
great variability of all the bones except those of the extremities. To a
certain extent we can understand why the skeleton fluctuates so much in
structure; fowls have been exposed to unnatural conditions of life, and
their whole organisation has thus been rendered variable; but the breeder
is quite indifferent to, and never intentionally selects, any modifications
in the skeleton. External characters, if not attended to by man,--such as
the number of the tail and wing feathers and their relative lengths, which
in wild birds are generally constant points,--fluctuate in our domestic
fowls in the same manner as the several parts of the skeleton. An
additional toe is a "point" in Dorkings, and has become a fixed character,
but is variable in {270} Cochins and Silk-fowls. The colour of the plumage
and the form of the comb are in most breeds, or even sub-breeds, eminently
fixed characters; but in Dorkings these points have not been attended to,
and are variable. When any modification in the skeleton is related to some
external character which man values, it has been, unintentionally on his
part, acted on by selection, and has become more or less fixed. We see this
in the wonderful protuberance of the skull, which supports the crest of
feathers in Polish fowls, and which by correlation has affected other parts
of the skull. We see the same result in the two protuberances which support
the horns in the horned fowl, and in the flattened shape of the front of
the skull in Hamburghs consequent on their flattened and broad
"rose-combs." We know not in the least whether additional ribs, or the
changed outline of the occipital foramen, or the changed form of the
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