Prince Bonaparte 'Coup d'oeil sur l'Ordre des Pigeons.'), and that this
reduction has acted by correlation on the beaks of the great majority of
the birds in Table 5.I. When, on the other hand, the beak has been much
elongated by the continued selection of successive slight increments of
length, the feet by correlation have likewise become much elongated in
comparison with those of the wild rock-pigeon, notwithstanding their
lessened use.
As I had taken measures from the end of the middle toe to the heel of the
tarsus in the rock-pigeon and in the above thirty-six birds, I have made
calculations analogous with those above given, and the result is the same--
namely, that in the short-beaked breeds, with equally few exceptions as in
the former case, the middle toe conjointly with the tarsus has decreased in
length; whereas in the long-beaked breeds it has increased in length,
though not quite so uniformly as in the former case, for the leg, in some
varieties of the Runt varies much in length.
LENGTH OF STERNUM.
As fancy pigeons are generally confined in aviaries of moderate size, and
as even when not confined they do not search for their own food, they must
during many generations have used their wings incomparably less than the
wild rock-pigeon. Hence it seemed to me probable that all the parts of the
skeleton subservient to flight would be found to be reduced in size. With
respect to the sternum, I have carefully measured its extreme length in
twelve birds of different breeds, and in two wild rock-pigeons from the
Shetland Islands. For the proportional comparison I have tried three
standards of measurement, with all twelve birds namely, the length from the
base of the beak to the oil-gland, to the end of the tail, and from the
extreme tip to tip of wings. The result has been in each case nearly the
same, the sternum being invariably found to be shorter than in the wild
rock-pigeon. I will give only a single table, as calculated by the standard
from the base of the beak to the oil-gland; for the result in this case is
nearly the mean between the results obtained by the two other standards.
TABLE 5.III.
LENGTH OF STERNUM.
Column 1. Name of Breed.
Column 2. Actual Length in Inches.
Column 3. Too short by (inches).
1. 2. 3.
Wild Rock-pigeon. 2.55
Pied Scanderoon. 2.80 0.60
Bagadotten Carrier. 2.80 0.17
Dragon. 2.45 0.41
Carrier. 2.75 0.35
Short-faced Tumbler. 2.05 0.28
Barb. 2.35 0.34
Nun. 2.27 0.15
German Pouter. 2.36 0.54
Jacobin. 2.33 0.22
English Frill-back. 2.40 0.43
Swallow. 2.45 0.17
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