We thus clearly see that the sternum, scapula, and furculum are all reduced
in proportional length; but when we turn to the wings we find what at first
appears a wholly different and unexpected result. I may here remark that I
have not picked out specimens, but have used every measurement made by me.
Taking the length from the base of beak to the end of the tail as the
standard of comparison, I find that, out of thirty-five birds of various
breeds, twenty-five have wings of greater, and ten have them of less
proportional length, than in the rock-pigeon. But from the frequently
correlated length of the tail and wing-feathers, it is better to take as
the standard of comparison the length from the base of the beak to the oil-
gland; and by this standard, out of twenty-six of the same birds which had
been thus measured, twenty-one had wings too long, and only five had them
too short. In the twenty-one birds the wings exceeded in length those of
the rock-pigeon, on an average, by 1 1/3 inch; whilst in the five birds
they were less in length by only .8 of an inch. As I was much surprised
that the wings of closely confined birds should thus so frequently have
been increased in length, it occurred to me that it might be solely due to
the greater length of the wing-feathers; for this certainly is the case
with the Jacobin, which has wings of unusual length. As in almost every
case I had measured the folded wings, I subtracted the length of this
terminal part from that of the expanded wings, and thus I obtained, with a
moderate degree of accuracy, the length of the wings from the ends of the
two radii, answering from wrist to wrist in our arms. The wings, thus
measured in the same twenty-five birds, now gave a widely different result;
for they were proportionally with those of the rock-pigeon too short in
seventeen birds, and in only eight too long. Of these eight birds, five
were long-beaked (5/39. It perhaps deserves notice that besides these five
birds two of the eight were Barbs, which, as I have shown, must be classed
in the same group with the long-beaked Carriers and Runts. Barbs may
properly be called short-beaked Carriers. It would, therefore, appear as
if, during the reduction of their beaks, their wings had retained a little
of that excess of length which is characteristic of their nearest relations
and progenitors.), and this fact perhaps indicates that there is some
correlation of the length of the beak with the length of the bones of the
wings, in the same manner as with that of the feet and tarsi. The
shortening of the humerus and radius in the seventeen birds may probably be
attributed to disuse, as in the case of the scapula and furculum to which
the wing-bones are attached;--the lengthening of the wing-feathers, and
consequently the expansion of the wings from tip to tip, being, on the
other hand, as completely independent of use and disuse as is the growth of
the hair or wool on our long-haired dogs or long-woolled sheep.
Public-domain text, read in full here on John Shaqi.
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