[Firstly, the Columba leuconota resembles certain domestic varieties in its
plumage, with the one marked and never-failing difference of a white band
which crosses the tail at some distance from the extremity. This species,
moreover, inhabits the Himalaya, close to the limit of perpetual snow; and
therefore, as Mr. Blyth has remarked, is not likely to have been the parent
of our domestic breeds, which thrive in the hottest countries. Secondly,
the C. rupestris, of Central Asia, which is intermediate (6/7. 'Proc.
Zoolog. Soc.' 1859 page 400.) between the C. leuconota and livia; but has
nearly the same coloured tail as the former species. Thirdly, the Columba
littoralis builds and roosts, according to Temminck, on rocks in the
Malayan archipelago; it is white, excepting parts of the wing and the tip
of the tail, which are black; its legs are livid-coloured, and this is a
character not observed in any adult domestic pigeon; but I need not have
mentioned this species or the closely-allied C. luctuosa, as they in fact
belong to the genus Carpophaga. Fourthly, Columba guinea, which ranges from
Guinea (6/8. Temminck 'Hist. Nat. Gen. des Pigeons' tome 1; also 'Les
Pigeons' par Mme. Knip and Temminck. Bonaparte, however, in his 'Coup-
d'oeil' believes that two closely allied species are confounded together
under this name. The C. leucocephala of the West Indies is stated by
Temminck to be a rock-pigeon; but I am informed by Mr. Gosse that this is
an error.) to the Cape of Good Hope, and roosts either on trees or rocks,
according to the nature of the country. This species belongs to the genus
Strictoenas of Reichenbach, but is closely allied to Columba; it is to some
extent coloured like certain domestic races, and has been said to be
domesticated in Abyssinia; but Mr. Mansfield Parkyns, who collected the
birds of that country and knows the species, informs me that this is a
mistake. Moreover, the C. guinea is characterised by the feathers of the
neck having peculiar notched tips,--a character not observed in any
domestic race. Fifthly, the Columba oenas of Europe, which roosts on trees,
and builds its nest in holes, either in trees or the ground; this species,
as far as external characters go, might be the parent of several domestic
races; but, though it crosses readily with the true rock-pigeon, the
offspring, as we shall presently see, are sterile hybrids, and of such
sterility there is not a trace when the domestic races are intercrossed. It
should also be observed that if we were to admit, against all probability,
that any of the foregoing five or six species were the parents of some of
our domestic pigeons, not the least light would be thrown on the chief
differences between the eleven most strongly-marked races.
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