Judging from the differences which generally distinguish the sexes in the
Gallinaceae, certain characters in our domestic fowls appear to have been
transferred from the one sex to the other. In all the species (except in
Turnix), when there is any conspicuous difference in plumage between the
male and female, the male is always the most beautiful; but in golden-
spangled Hamburghs the hen is equally beautiful with the cock, and
incomparably more beautiful than the hen in any natural species of Gallus;
so that here a masculine character has been transferred to the female. On
the other hand, in Cuckoo Dorkings and in other cuckoo breeds the
pencilling, which in Gallus is a female attribute, has been transferred to
the male: nor, on the principle of analogous variation, is this
transference surprising, as the males in many gallinaceous genera are
barred or pencilled. With most of these birds head ornaments of all kinds
are more fully developed in the male than in the female; but in Polish
fowls the crest or top-knot, which in the male replaces the comb, is
equally developed in both sexes. In the males of certain other sub-breeds,
which from the hen having a small crest, are called lark-crested, "a single
upright comb sometimes almost entirely takes the place of the crest."
(7/54. Dixon 'Ornamental and Domestic Poultry' page 320.) From this latter
case, and more especially from some facts presently to be given with
respect to the protuberance of the skull in Polish fowls, the crest in this
breed must be viewed as a feminine character which has been transferred to
the male. In the Spanish breed the male, as we know, has an immense comb,
and this has been partially transferred to the female, for her comb is
unusually large, though not upright. In Game fowls the bold and savage
disposition of the male has likewise been largely transferred to the female
(7/55. Mr. Tegetmeier informs me that Game hens have been found so
combative, that it is now generally the practice to exhibit each hen in a
separate pen.); and she sometimes even possesses the eminently masculine
character of spurs. Many cases are on record of fertile hens being
furnished with spurs; and in Germany, according to Bechstein (7/56.
'Naturgeschichte Deutschlands' b. 3 1793 s. 339, 407.), the spurs in the
Silk hen are sometimes very long. He mentions also another breed similarly
characterised, in which the hens are excellent layers, but are apt to
disturb and break their eggs owing to their spurs.
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