an infinite number of the finest gradations, sometimes in one character and
sometimes in another, but all leading towards our present perfect type. By
small and doubtful steps such as these, nature, as we may confidently believe,
has progressed, on her grand march of improvement and development.
A similar line of reasoning is as applicable to separate organs as to the
whole organisation. A writer (20/91. Mr. J.J. Murphy in his opening address to
the Belfast Nat. Hist. Soc. as given in the 'Belfast Northern Whig' November
19, 1866. Mr. Murphy here follows the line of argument against my views
previously and more cautiously given by the Rev. C. Pritchard, Pres. Royal
Astronomical Soc., in his sermon Appendix page 33 preached before the British
Association at Nottingham 1866.) has recently maintained that "it is probably
no exaggeration to suppose that in order to improve such an organ as the eye
at all, it must be improved in ten different ways at once. And the
improbability of any complex organ being produced and brought to perfection in
any such way is an improbability of the same kind and degree as that of
producing a poem or a mathematical demonstration by throwing letters at random
on a table." If the eye were abruptly and greatly modified, no doubt many
parts would have to be simultaneously altered, in order that the organ should
remain serviceable.
But is this the case with smaller changes? There are persons who can see
distinctly only in a dull light, and this condition depends, I believe, on the
abnormal sensitiveness of the retina, and is known to be inherited. Now if a
bird, for instance, receive some great advantage from seeing well in the
twilight, all the individuals with the most sensitive retina would succeed
best and be the most likely to survive; and why should not all those which
happened to have the eye itself a little larger, or the pupil capable of
greater dilatation, be likewise preserved, whether or not these modifications
were strictly simultaneous? These individuals would subsequently intercross
and blend their respective advantages. By such slight successive changes, the
eye of a diurnal bird would be brought into the condition of that of an owl,
which has often been advanced as an excellent instance of adaptation. Short-
sight, which is often inherited, permits a person to see distinctly a minute
object at so near a distance that it would be indistinct to ordinary eyes; and
here we have a capacity which might be serviceable under certain conditions,
abruptly gained. The Fuegians on board the Beagle could certainly see distant
objects more distinctly than our sailors with all their long practice; I do
not know whether this depends upon sensitiveness or on the power of adjustment
in the focus; but this capacity for distant vision might, it is probable, be
slightly augmented by successive modifications of either kind. Amphibious
animals which are enabled to see both in the water and in the air, require and
possess, as M.
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