The importance of selection is likewise shown by considering special
characters. For instance, with most breeds of fowls the form of the comb and
the colour of the plumage have been attended to, and are eminently
characteristic of each race; but in Dorkings fashion has never demanded
uniformity of comb or colour; and the utmost diversity in these respects
prevails. Rose-combs, double-combs, cup-combs, etc., and colours of all kinds,
may be seen in purely bred and closely related Dorking fowls, whilst other
points, such as the general form of body, and the presence of an additional
toe, have been attended to, and are invariably present. It has also been
ascertained that colour can be fixed in this breed, as well as in any other.
(21/54. Mr. Baily in 'The Poultry Chronicle' volume 2 1854 page 150. Also
volume 1 page 342; volume 3 page 245.)
During the formation or improvement of a breed, its members will always be
found to vary much in those characters to which especial attention is
directed, and of which each slight improvement is eagerly sought and selected.
Thus, with short-faced tumbler-pigeons, the shortness of the beak, shape of
head and plumage,--with carriers, the length of the beak and wattle,--with
fantails, the tail and carriage,--with Spanish fowls, the white face and
comb,--with long-eared rabbits, the length of ear, are all points which are
eminently variable. So it is in every case; and the large price paid for
first-rate animals proves the difficulty of breeding them up to the highest
standard of excellence. This subject has been discussed by fanciers (21/55.
'Cottage Gardener' 1855 December page 171; 1856 January pages 248, 323.), and
the greater prizes given for highly improved breeds, in comparison with those
given for old breeds which are not now undergoing rapid improvement, have been
fully justified. Nathusius makes (21/56. 'Ueber Shorthorn Rindvieh' 1857 s.
51.) a similar remark when discussing the less uniform character of improved
Shorthorn cattle and of the English horse, in comparison, for example, with
the unennobled cattle of Hungary, or with the horses of the Asiatic steppes.
This want of uniformity in the parts which at the time are undergoing
selection chiefly depends on the strength of the principle of reversion; but
it likewise depends to a certain extent on the continued variability of the
parts which have recently varied. That the same parts do continue varying in
the same manner we must admit, for if it were not so, there could be no
improvement beyond an early standard of excellence, and we know that such
improvement is not only possible, but is of general occurrence.
Public-domain text, read in full here on John Shaqi.
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