In a few instances selection, guided by utility for a single purpose, has led
to convergence of character. All the improved and different races of the pig,
as Nathusius has well shown (21/59. 'Die Racen des Schweines' 1860 s. 48.),
closely approach each other in character, in their shortened legs and muzzles,
their almost hairless, large, rounded bodies, and small tusks. We see some
degree of convergence in the similar outline of the body in well-bred cattle
belonging to distinct races. (21/60. See some good remarks on this head by M.
de Quatrefages 'Unite de l'Espece Humaine' 1861 page 119.) I know of no other
such cases.
Continued divergence of character depends on, and is indeed a clear proof, as
previously remarked, of the same parts continuing to vary in the same
direction. The tendency to mere general variability or plasticity of
organisation can certainly be inherited, even from one parent, as has been
shown by Gartner and Kolreuter, in the production of varying hybrids from two
species, of which one alone was variable. It is in itself probable that, when
an organ has varied in any manner, it will again vary in the same manner, if
the conditions which first caused the being to vary remain, as far as can be
judged, the same. This is either tacitly or expressly admitted by all
horticulturists: if a gardener observes one or two additional petals in a
flower, he feels confident that in a few generations he will be able to raise
a double flower, crowded with petals. Some of the seedlings from the weeping
Moccas oak were so prostrate that they only crawled along the ground. A
seedling from the fastigiate or upright Irish yew is described as differing
greatly from the parent-form "by the exaggeration of the fastigiate habit of
its branches." (21/61. Verlot 'Des Varietes' 1865 page 94.) Mr. Shirreff, who
has been highly successful in raising new kinds of wheat, remarks, "A good
variety may safely be regarded as the forerunner of a better one." (21/62. Mr.
Patrick Shirreff 'Gardener's Chronicle' 1858 page 771.) A great rose-grower,
Mr. Rivers, has made the same remark with respect to roses. Sageret (21/63.
'Pomologie Physiolog.' 1830 page 106.), who had large experience, in speaking
of the future progress of fruit-trees, observes that the most important
principle is "that the more plants have departed from their original type, the
more they tend to depart from it." There is apparently much truth in this
remark; for we can in no other way understand the surprising amount of
difference between varieties in the parts or qualities which are valued,
whilst other parts retain nearly their original character.
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