[Notwithstanding the facts previously given on the occurrence of highly
peculiar local diseases and on the strange modifications of structure in
plants caused by the inoculated poison of insects, and other analogous cases;
still there are a multitude of variations--such as the modified skull of the
niata ox and bulldog, the long horns of Caffre cattle, the conjoined toes of
the solid-hoofed swine, the immense crest and protuberant skull of Polish
fowls, the crop of the pouter-pigeon, and a host of other such cases--which we
can hardly attribute to the definite action, in the sense before specified, of
the external conditions of life. No doubt in every case there must have been
some exciting cause; but as we see innumerable individuals exposed to nearly
the same conditions, and one alone is affected, we may conclude that the
constitution of the individual is of far higher importance than the conditions
to which it has been exposed. It seems, indeed, to be a general rule that
conspicuous variations occur rarely, and in one individual alone out of
millions, though all may have been exposed, as far as we can judge, to nearly
the same conditions. As the most strongly marked variations graduate
insensibly into the most trifling, we are led by the same train of thought to
attribute each slight variation much more to innate differences of
constitution, however caused, than to the definite action of the surrounding
conditions.
We are led to the same conclusion by considering the cases, formerly alluded
to, of fowls and pigeons, which have varied and will no doubt go on varying in
directly opposite ways, though kept during many generations under nearly the
same conditions. Some, for instance, are born with their beaks, wings, tails,
legs, etc., a little longer, and others with these same parts a little
shorter. By the long-continued selection of such slight individual differences
which occur in birds kept in the same aviary, widely different races could
certainly be formed; and long-continued selection, important as is the result,
does nothing but preserve the variations which arise, as it appears to us,
spontaneously.
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