The organisation or constitution of the being which is acted on, is generally
a much more important element than the nature of the changed conditions, in
determining the nature of the variation. We have evidence of this in the
appearance of nearly similar modifications under different conditions, and of
different modifications under apparently nearly the same conditions. We have
still better evidence of this in closely parallel varieties being frequently
produced from distinct races, or even distinct species; and in the frequent
recurrence of the same monstrosity in the same species. We have also seen that
the degree to which domesticated birds have varied, does not stand in any
close relation with the amount of change to which they have been subjected.
To recur once again to bud-variations. When we reflect on the millions of buds
which many trees have produced, before some one bud has varied, we are lost in
wonder as to what the precise cause of each variation can be. Let us recall
the case given by Andrew Knight of the forty-year-old tree of the yellow
magnum bonum plum, an old variety which has been propagated by grafts on
various stocks for a very long period throughout Europe and North America, and
on which a single bud suddenly produced the red magnum bonum. We should also
bear in mind that distinct varieties, and even distinct species,--as in the
case of peaches, nectarines, and apricots,--of certain roses and camellias,--
although separated by a vast number of generations from any progenitor in
common, and although cultivated under diversified conditions, have yielded by
bud-variation closely analogous varieties. When we reflect on these facts we
become deeply impressed with the conviction that in such cases the nature of
the variation depends but little on the conditions to which the plant has been
exposed, and not in any especial manner on its individual character, but much
more on the inherited nature or constitution of the whole group of allied
beings to which the plant in question belongs. We are thus driven to conclude
that in most cases the conditions of life play a subordinate part in causing
any particular modification; like that which a spark plays, when a mass of
combustibles bursts into flame--the nature of the flame depending on the
combustible matter, and not on the spark. (23/63. Professor Weismann argues
strongly in favour of this view in his 'Saison-Dimorphismus der
Schmetterlinge' 1875 pages 40-43.)
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