pure breeds, written as it were in invisible ink, yet ready at any time to be
evolved under certain conditions.
What these conditions precisely are, we do not know. But any cause which
disturbs the organisation or constitution seems to be sufficient. A cross
certainly gives a strong tendency to the reappearance of long-lost characters,
both corporeal and mental. In the case of plants, this tendency is much
stronger with those species which have been crossed after long cultivation and
which therefore have had their constitutions disturbed by this cause as well
as by crossing, than with species which have always lived under their natural
conditions and have then been crossed. A return, also, of domesticated animals
and cultivated plants to a wild state favours reversion; but the tendency
under these circumstances has been much exaggerated.
When individuals of the same family which differ somewhat, and when races or
species are crossed, the one is often prepotent over the other in transmitting
its character. A race may possess a strong power of inheritance, and yet when
crossed, as we have seen with trumpeter-pigeons, yield to the prepotency of
every other race. Prepotency of transmission may be equal in the two sexes of
the same species, but often runs more strongly in one sex. It plays an
important part in determining the rate at which one race can be modified or
wholly absorbed by repeated crosses with another. We can seldom tell what
makes one race or species prepotent over another; but it sometimes depends on
the same character being present and visible in one parent, and latent or
potentially present in the other.
Characters may first appear in either sex, but oftener in the male than in the
female, and afterwards be transmitted to the offspring of the same sex. In
this case we may feel confident that the peculiarity in question is really
present though latent in the opposite sex! hence the father may transmit
through his daughter any character to his grandson; and the mother conversely
to her granddaughter. We thus learn, and the fact is an important one, that
transmission and development are distinct powers. Occasionally these two
powers seem to be antagonistic, or incapable of combination in the same
individual; for several cases have been recorded in which the son has not
directly inherited a character from his father, or directly transmitted it to
his son, but has received it by transmission through his non-affected mother,
and transmitted it through his non-affected daughter. Owing to inheritance
being limited by sex, we see how secondary sexual characters may have arisen
under nature; their preservation and accumulation being dependent on their
service to either sex.
Public-domain text, read in full here on John Shaqi.
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