It is remarkable, as has been strongly insisted upon by Isidore Geoffroy St.
Hilaire in regard to animals, that the transmission of characters without
fusion occurs very rarely when species are crossed; I know of one exception
alone, namely, with the hybrids naturally produced between the common and
hooded crow (Corvus corone and cornix), which, however, are closely allied
species, differing in nothing except colour. Nor have I met with any well-
ascertained cases of transmission of this kind, even when one form is strongly
prepotent over another, when two races are crossed which have been slowly
formed by man's selection, and therefore resemble to a certain extent natural
species. Such cases as puppies in the same litter closely resembling two
distinct breeds, are probably due to superfoetation,--that is, to the
influence of two fathers. All the characters above enumerated, which are
transmitted in a perfect state to some of the offspring and not to others,--
such as distinct colours, nakedness of skin, smoothness of leaves, absence of
horns or tail, additional toes, pelorism, dwarfed structure, etc.,--have all
been known to appear suddenly in individual animals and plants. From this
fact, and from the several slight, aggregated differences which distinguish
domestic races and species from one another, not being liable to this peculiar
form of transmission, we may conclude that it is in some way connected with
the sudden appearance of the characters in question.]
ON THE MODIFICATION OF OLD RACES AND THE FORMATION OF NEW RACES BY CROSSING.
We have hitherto chiefly considered the effects of crossing in giving
uniformity of character; we must now look to an opposite result. There can be
no doubt that crossing, with the aid of rigorous selection during several
generations, has been a potent means in modifying old races, and in forming
new ones. Lord Orford crossed his famous stud of greyhounds once with the
bulldog, in order to give them courage and perseverance. Certain pointers have
been crossed, as I hear from the Rev. W.D. Fox, with the foxhound, to give
them dash and speed. Certain strains of Dorking fowls have had a slight
infusion of Game blood; and I have known a great fancier who on a single
occasion crossed his turbit-pigeons with barbs, for the sake of gaining
greater breadth of beak.
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