Before giving the few recorded cases of forms, which must be ranked as
varieties, being in some degree sterile when crossed, I may remark that other
causes sometimes interfere with varieties freely intercrossing. Thus they may
differ too greatly in size, as with some kinds of dogs and fowls: for
instance, the editor of the 'Journal of Horticulture, etc.' (16/2. December
1863 page 484.) says that he can keep Bantams with the larger breeds without
much danger of their crossing, but not with the smaller breeds, such as Games,
Hamburghs, etc. With plants a difference in the period of flowering serves to
keep varieties distinct, as with the various kinds of maize and wheat: thus
Colonel Le Couteur (16/3. On 'The Varieties of Wheat' page 66.) remarks, "the
Talavera wheat, from flowering much earlier than any other kind, is sure to
continue pure." In different parts of the Falkland Islands the cattle are
breaking up into herds of different colours; and those on the higher ground,
which are generally white, usually breed, as I am informed by Sir J. Sulivan,
three months earlier than those on the lowland; and this would manifestly tend
to keep the herds from blending.
Certain domestic races seem to prefer breeding with their own kind; and this
is a fact of some importance, for it is a step towards that instinctive
feeling which helps to keep closely allied species in a state of nature
distinct. We have now abundant evidence that, if it were not for this feeling,
many more hybrids would be naturally produced than in this case. We have seen
in the first chapter that the alco dog of Mexico dislikes dogs of other
breeds; and the hairless dog of Paraguay mixes less readily with the European
races, than the latter do with each other. In Germany the female Spitz-dog is
said to receive the fox more readily than will other dogs; a female Australian
Dingo in England attracted the wild male foxes. But these differences in the
sexual instinct and attractive power of the various breeds may be wholly due
to their descent from distinct species. In Paraguay the horses have much
freedom, and an excellent observer (16/4. Rengger 'Saugethiere von Paraguay'
s. 336.) believes that the native horses of the same colour and size prefer
associating with each other, and that the horses which have been imported from
Entre Rios and Banda Oriental into Paraguay likewise prefer associating
together. In Circassia six sub-races of the horse have received distinct
names; and a native proprietor of rank (16/5. See a memoir by MM. Lherbette
and De Quatrefages in 'Bull. Soc. d'Acclimat.' tome 8 July 1861 page 312.)
asserts that horses of three of these races, whilst living a free life, almost
always refuse to mingle and cross, and will even attack one another.
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