_Effects of Selection, as shown by the parts most valued by man presenting
the greatest amount of Difference._--The power of long-continued selection,
whether methodical or unconscious, or both combined, is well shown in a
general way, namely, by the comparison of the differences between the
varieties of distinct species, which are valued for different parts, such
as for the leaves, or stems, or tubers, the seed, or fruit, or flowers.
Whatever part man values most, that part will be found to present the
greatest amount of difference. With trees cultivated for their fruit,
Sageret remarks that the fruit is larger than in the parent-species, whilst
with those cultivated for the seed, as with nuts, walnuts, almonds,
chesnuts, &c., it is the seed itself which is larger; and he accounts for
this fact by the fruit in the one case, and by the seed in the other,
having been carefully attended to and selected during many ages. Gallesio
has made the same observation. Godron insists on the diversity of the tuber
in the potato, of the bulb in the onion, and of the fruit in the melon; and
on the close similarity in these same plants of the other parts.[529]
In order to judge how far my own impression on this subject was correct, I
cultivated numerous varieties of the same species close to each other. The
comparison of the amount of difference between widely different organs is
necessarily vague; I will therefore give the results in only a few cases.
We have previously seen in the ninth chapter how greatly the varieties of
the cabbage differ in their foliage and stems, which are the selected
parts, and how closely they resembled each other in their flowers,
capsules, and seeds. In seven varieties of the radish, the roots differed
greatly in colour and shape, but no difference {218} whatever could be
detected in their foliage, flowers, or seeds. Now what a contrast is
presented, if we compare the flowers of the varieties of these two plants
with those of any species cultivated in our flower-gardens for ornament; or
if we compare their seeds with those of the varieties of maize, peas,
beans, &c., which are valued and cultivated for their seeds. In the ninth
chapter it was shown that the varieties of the pea differ but little except
in the tallness of the plant, moderately in the shape of the pod, and
greatly in the pea itself, and these are all selected points. The
varieties, however, of the _Pois sans parchemin_ differ much more in their
pods, and these are eaten and valued. I cultivated twelve varieties of the
common bean; one alone, the Dwarf Fan, differed considerably in general
appearance; two differed in the colour of their flowers, one being an
albino, and the other being wholly instead of partially purple; several
differed considerably in the shape and size of the pod, but far more in the
bean itself, and this is the valued and selected part. Toker's bean, for
instance, is twice-and-a-half as long and broad as the horse-bean, and is
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