Of all the causes which induce variability, excess of food, whether or not
changed in nature, is probably the most powerful. This view was held with
regard to plants by Andrew Knight, and is now held by Schleiden, more
especially in reference to the inorganic elements of the food.[612] In
order to give a plant more food it suffices in most cases to grow it
separately, and thus prevent other plants robbing its roots. It is
surprising, as I have often seen, how vigorously our common wild plants
flourish when planted by themselves, though not in highly manured land.
Growing plants separately is, in fact, the first step in cultivation. We
see the converse of the belief that excess of food induces variability in
the following statement by a great raiser of seeds of all kinds.[613] "It
is a rule invariably with us, when we desire to keep a true stock of any
one kind of seed, to grow it on poor land without dung; but when we grow
for quantity, we act contrary, and sometimes have dearly to repent of it."
In the case of animals the want of a proper amount of exercise, as
Bechstein has remarked, has perhaps played, independently of the direct
effects of the disuse of any particular organ, an important part in causing
variability. We can see in a vague manner that, when the organised and
nutrient fluids of the body are not used during growth, or by the wear and
tear of the tissues, {258} they will be in excess; and as growth,
nutrition, and reproduction are intimately allied processes, this
superfluity might disturb the due and proper action of the reproductive
organs, and consequently affect the character of the future offspring. But
it may be argued that neither an excess of food nor a superfluity in the
organised fluids of the body necessarily induces variability. The goose and
the turkey have been well fed for many generations, yet have varied very
little. Our fruit-trees and culinary plants, which are so variable, have
been cultivated from an ancient period, and, though they probably still
receive more nutriment than in their natural state, yet they must have
received during many generations nearly the same amount; and it might be
thought that they would have become habituated to the excess. Nevertheless,
on the whole, Knight's view, that excess of food is one of the most potent
causes of variability, appears, as far as I can judge, probable.
Whether or not our various cultivated plants have received nutriment in
excess, all have been exposed to changes of various kinds. Fruit-trees are
grafted on different stocks, and grown in various soils. The seeds of
culinary and agricultural plants are carried from place to place; and
during the last century the rotation of our crops and the manures used have
been greatly changed.
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