Some flowers have almost certainly become more or less completely peloric through reversion. _Corydalis tuberosa_ properly has one of its two nectaries colourless, destitute of nectar, only half the size of the other, and {59} therefore, to a certain extent, in a rudimentary state; the pistil is curved towards the perfect nectary, and the hood, formed of the inner petals, slips off the pistil and stamens in one direction alone, so that, when a bee sucks the perfect nectary, the stigma and stamens are exposed and rubbed against the insect's body. In several closely allied genera, as in Dielytra, &c., there are two perfect nectaries, the pistil is straight, and the hood slips off on either side, according as the bee sucks either nectary. Now, I have examined several flowers of _Corydalis tuberosa_, in which both nectaries were equally developed and contained nectar; in this we see only the redevelopment of a partially aborted organ; but with this redevelopment the pistil becomes straight, and the hood slips off in either direction; so that these flowers have acquired the perfect structure, so well adapted for insect agency, of Dielytra and its allies. We cannot attribute these coadapted modifications to chance, or to correlated variability; we must attribute them to reversion to a primordial condition of the species. The peloric flowers of Pelargonium have their five petals in all respects alike, and there is no nectary; so that they resemble the symmetrical flowers of the closely allied Geranium-genus; but the alternate stamens are also sometimes destitute of anthers, the shortened filaments being left as rudiments, and in this respect they resemble the symmetrical flowers of the closely allied genus, Erodium. Hence we are led to look at the peloric flowers of Pelargonium as having probably reverted to the state of some primordial form, the progenitor of the three closely related genera of Pelargonium, Geranium, and Erodium.