_On certain Characters not blending._--When two breeds are crossed
their characters usually become intimately fused together; but some
characters refuse to blend, and are transmitted in an unmodified state
either from both parents or from one. When grey and white mice are
paired, the young are not piebald nor of an intermediate tint, but are
pure white or of the ordinary grey colour: so it is when white and
common collared turtle-doves are paired. In breeding Game fowls, a
great authority, Mr. J. Douglas, remarks, "I may here state a strange
fact: if you cross a black with a white game, you get birds of both
breeds of the clearest colour." Sir R. Heron crossed during many years
white, black, brown, and fawn-coloured Angora rabbits, and never once
got these colours mingled in the same animal, but often all four
colours in the same litter.[196] Additional cases could be given, but
this form of inheritance is very far from universal even with respect
to the most distinct colours. When turnspit dogs and ancon sheep, both
of which have dwarfed limbs, are crossed with common breeds, the
offspring are not intermediate in structure, but take after either
parent. When tailless or hornless animals are crossed with perfect
animals, it frequently, but by no means invariably, happens that the
offspring are {93} either perfectly furnished with these organs or are
quite destitute of them. According to Rengger, the hairless condition
of the Paraguay dog is either perfectly or not at all transmitted to
its mongrel offspring; but I have seen one partial exception in a dog
of this parentage which had part of its skin hairy, and part naked; the
parts being distinctly separated as in a piebald animal. When Dorking
fowls with five toes are crossed with other breeds, the chickens often
have five toes on one foot and four on the other. Some crossed pigs
raised by Sir R. Heron between the solid-hoofed and common pig had not
all four feet in an intermediate condition, but two feet were furnished
with properly divided, and two with united hoofs.
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