The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
In the shoulder-girdle the coracoids are large and meet in a ventral
symphysis; precoracoids and a sternum are apparently absent, but parts
generally regarded[71] as the clavicles and interclavicle are well
developed. In the pelvis, the pubes and ischia meet in a long
symphysis. The limbs are pentedactylate, and in the best known forms,
the Plesiosauridae, form swimming paddles.
The Sauropterygia occur in beds of Secondary age, and some of the best
known genera are _Plesiosaurus_, _Pliosaurus_ and _Nothosaurus_.
_Order 3._ CHELONIA.
In the Tortoises and Turtles the body is enclosed in a bony box,
formed of the dorsal carapace, and a flat ventral buckler, the
plastron. Except in _Dermochelys_ the carapace is partly formed from
the vertebral column and ribs, partly from dermal bones. Both carapace
and plastron are, except in _Dermochelys_, _Trionyx_ and their allies,
covered with an epidermal exoskeleton of horny plates, which are
regularly arranged, though their outlines do not coincide with those
of the underlying bones. The thoracic vertebrae have no transverse
processes, and are quite immovably fixed, but the cervical and caudal
vertebrae are very freely movable. There are no lumbar vertebrae. The
skull is extremely solid, and frequently has a very complete false
roof. Teeth have been detected in embryos of _Trionyx_ but with this
exception the jaws are toothless, and are encased in horny beaks. The
quadrate is firmly fixed. The facial part of the skull is very short,
and the alisphenoidal and orbitosphenoidal regions are unossified. In
living forms there are no separate nasal bones, while large
prefrontals and postfrontals are developed. There is a comparatively
complete bony palate chiefly formed of the palatines and pterygoids.
The anterior nares are united and placed at the anterior end of the
skull, and the premaxillae are very small. There is no transpalatine
bone and the vomer is unpaired. The dentaries are generally fused
together.
There are ten pairs of ribs, and each rib has only a single head and
is partially attached to two vertebrae; there are no cervical or
sternal ribs. There is no true sternum.
The three anterior elements of the plastron are respectively
homologous with the interclavicle and two clavicles of other reptiles,
while the remaining elements of the plastron are probably homologous
with the abdominal ribs of Crocodiles. The pectoral girdle lies within
the ribs, and the precoracoids and coracoids do not meet in ventral
symphyses. The scapula and precoracoid are ossified continuously. The
pubis probably corresponds with the pre-pubis of Dinosaurs. There are
four limbs each with five digits.
The order includes three suborders:--
_Suborder (1)._ TRIONYCHIA.
The carapace and plastron have a rough granular surface covered with
skin and without any horny shields.
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