The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
The teeth are, in the adult, planted in separate deep sockets. The
skull is very dense and solid, and all the component bones including
the quadrate are firmly united. The dorsal surface of the skull is
generally characteristically sculptured. There is an interorbital
septum, and the orbitosphenoidal and presphenoidal regions are
imperfectly ossified. Supratemporal, infratemporal, and post-temporal
fossae occur, but no interparietal foramen. In living genera there is
a long secondary palate formed by the meeting in the middle line of
the palatines, pterygoids and maxillae (fig. 43, A).
Cervical ribs (fig. 41, 8 and 9) are well developed, and articulate
with rather prominent surfaces borne on the neural arches and centra
respectively. The thoracic ribs articulate with the long transverse
processes, and sternal ribs and abdominal splint ribs (fig. 46, 4)
occur. The sternum is cartilaginous, and both it and the
shoulder-girdle are very simple. The precoracoid is represented by
merely a small process on the coracoid, while the clavicles are
absent, except in the Parasuchia. In the pelvis (fig. 49) there is a
large ilium, and an ischium meeting its fellow in a ventral symphysis;
these two bones form almost the whole of the acetabulum. In front of
the acetabulum, in the Eusuchia, projects a bone which is generally
called the pubis, but is in reality rather an epipubis (fig. 49, 4),
the true pubis being probably represented by a fourth element which
remains cartilaginous for some time, and later on ossifies and
attaches itself to the ischium. The limbs are small in proportion to
the size of the body, and are adapted for swimming or for shuffling
along the ground; they are plantigrade and the bones are all solid. In
living forms the anterior limbs have five digits and the posterior
four, the fifth being represented only by a short metatarsal. The
first three digits in each case are clawed. The calcaneum has a large
backwardly-projecting process.
The order Crocodilia may be subdivided into two suborders.
_Suborder (1)._ PARASUCHIA.
The vertebral centra are flat or biconcave. The premaxillae are very
large, and the nares are separated, and placed far back. The posterior
narial openings lie comparatively far forward between the anterior
extremities of the palatines.
The palatines and pterygoids do not form a secondary palate. The
supratemporal fossae are small, and open posteriorly, the lateral
temporal fossae are very large. The parietals and frontals are paired.
Clavicles are present. The best known and most important genus of
these extinct crocodiles is _Belodon_.
_Suborder_ (2). EUSUCHIA.
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