The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
[88] See H.G. Seeley, On the Organisation of the Ornithosauria,
_Journ. Linn. Soc._ (Zoology) vol. XIII. p. 84. K.A. Zittel, Ueber
Flugsaurier aus dem lithographischen schiefer, _Palaeontograph._ XXIX.
p. 49.
CHAPTER XIV.
THE SKELETON OF THE GREEN TURTLE.
(_Chelone midas._)
The most striking feature as regards the skeleton of the Turtle is
that the trunk is enveloped in a bony box, the dorsal portion of which
is called the =carapace=, while the ventral portion is the =plastron=.
I. EXOSKELETON.
_a._ The =epidermal exoskeleton= in the Green Turtle as in all other
Chelonia except _Dermochelys_, _Trionyx_ and their allies is strongly
developed, its most important part consisting of a series of horny
=shields= which cover over the bony plates of the carapace and
plastron but do not at all correspond to them in size and arrangement.
The shields covering over the =carapace= consist of three rows of
larger central shields,--five (=vertebral=) shields being included in
the middle row and four (=costal=) in each lateral row,--and of a
number of smaller =marginal= shields.
Of the marginal shields, that lying immediately in front of the first
vertebral is termed the =nuchal=, while the two succeeding the last
vertebral are called sometimes =pygal=, sometimes =supracaudal=; the
remainder are the marginal shields proper.
The epidermal covering of the =plastron= consists principally of six
pairs of symmetrically arranged shields, called respectively the
=gular=, =humeral=, =pectoral=, =abdominal=, =femoral=, and =anal=,
the gular being the most anterior. In front of the gular shields is an
unpaired =intergular=, and the shields of the plastron are connected
laterally with those of the carapace, by five or six pairs of rather
irregular =infra-marginal= shields. Smaller horny plates occur on
other parts of the body, especially on the limbs and head.
Two other sets of structures belong also to the epidermal exoskeleton,
viz. (_a_) horny =beaks= with denticulated edges which ensheath both
upper and lower jaws, (_b_) =claws=, which as a rule are borne only by
the first digit of each limb. Sometimes in young individuals the
second digit is also clawed.
_b._ The =dermal exoskeleton= is strongly developed, and is combined
with endoskeletal structures derived from the ribs and vertebrae to
form the carapace.
The =Carapace= (fig. 36) consists of a number of plates firmly united
to one another by sutures. They have a very definite arrangement and
include:
(_a_) the =nuchal= plate (fig. 36, 1), a wide plate forming the whole
of the anterior margin of the carapace. It is succeeded by three
series of plates, eight in each series, which together make up the
main part of the carapace. Of these the small
(_b_) =neural plates=[1] (fig. 36, A, 2) form the middle series. They
are closely united with the neural arches of the underlying vertebrae;
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