The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
The _frontals_ are a pair of small bones lying immediately in front of
the parietals, and in front of them are the _prefrontals_ (figs. 38
and 39, 20), a pair of similar but still smaller bones, which are
produced ventrally to meet the vomer and palatines. They form also the
dorsal boundary of the anterior nares. The _postfrontals_ (figs. 38
and 39, 18) are larger bones, united dorsally to the frontals and
parietals, posteriorly to the squamosals, and ventrally to the jugals
and quadratojugals. All three pairs of frontal bones, especially the
postfrontals, take part in the bounding of the orbits.
[Illustration FIG. 39. LONGITUDINAL VERTICAL SECTION THROUGH THE
CRANIUM OF A GREEN TURTLE (_Chelone midas_). × 2/3. (Camb. Mus.)
1. parietal.
2. squamosal.
3. quadrate.
4. basisphenoid.
5. basi-occipital.
6. quadratojugal.
7. pro-otic.
8. opisthotic.
9. pterygoid.
10. palatine.
11. rod passed into narial passage.
12. exoccipital.
13. epi-otic fused to supra-occipital.
14. supra-occipital.
15. premaxillae.
16. maxillae.
17. jugal.
18. postfrontal.
19. vomer.
20. prefrontal.
21. frontal.
V, VII, VIII, IX, X, XI, XII,
foramina for the exit of
cranial nerves.]
2. THE SENSE CAPSULES.
Skeletal structures occur in connection with each of the three special
sense organs of hearing, sight, and smell.
The =Auditory capsules=.
The auditory or periotic capsule of the turtle is rather large and its
walls are well ossified, epi-otic, pro-otic and opisthotic bones being
present.
The =epi-otic= (fig. 39, 13) is the more dorsal of the three bones,
and in the adult is completely ankylosed with the supra-occipital.
The =opisthotic= (fig. 39, 8) is the ventral posterior element. On its
inner side it is united to the supra-occipital above, and to the
exoccipital below; it sometimes becomes completely fused with the
exoccipital. In front it meets the pro-otic, and on its outer side the
squamosal and quadrate. Its anterior portion is hollowed out by the
cavity in which the auditory organ lies, it gives off also a process
which is separated from the exoccipital by an oval foramen through
which the glossopharyngeal, pneumogastric, and spinal accessory nerves
leave the cranial cavity.
The =pro-otic= is the anterior element; it meets the supra-occipital
and opisthotic posteriorly, while anteriorly it is separated from the
alisphenoidal plate of the parietal and pterygoid by a large oval
foramen through which the maxillary and mandibular branches of the
trigeminal nerve pass out (fig. 39, V 1 & 2). It is hollowed out
posteriorly by the cavity in which the auditory organ lies, and its
inner wall as seen in longitudinal section is pierced by a foramen
through which the external carotid artery and facial nerve leave the
cranial cavity,--the nerve finally leaving the skull through a small
oval foramen on the anterior face of the pro-otic near its junction
with the quadrate.
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