The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
The =alisphenoids= (fig. 45, 13) are a pair of irregular bones which
arise from the basisphenoid antero-laterally, and are united dorsally
with the parietal, frontal, and postfrontals. They bound most of the
anterior part of the brain case, and each presents on its inner face a
deep concavity which lodges the cerebral hemisphere of its side.
Viewed from the ventral side the two alisphenoids are seen to almost
or quite meet one another immediately below the frontal, and then to
diverge, forming an irregular opening--partially closed by cartilage
in the fresh specimen,--through which the optic nerves leave the
cranial cavity. Further back the alisphenoids meet one another for a
narrow area, and then diverge again, so that between each and the
rostrum of the basisphenoid there appears an opening (fig. 44, III,
VI) through which the oculomotor and abducens nerves leave the
cranium. Further back still each is united for a short space with the
basisphenoid, pterygoid and quadrate, and then becomes separated from
the quadrate by a large foramen, the =foramen ovale= (fig. 44, V),
through which the whole of the trigeminal nerve passes out.
The dorsal portion of the parietal segment is formed by the _parietal_
(fig. 45, 4), which though double in the embryo, early comes to form a
single bone. It extends over the posterior part of the cranial cavity,
and is continuous in front with the frontal, behind with the
supra-occipital, and laterally with the postfrontals, squamosals,
alisphenoids, pro-otics and epi-otics. It forms the inner boundary of
a large rounded vacuity on the roof of the skull, the =supratemporal
fossa=.
The =frontal segment= is very imperfectly ossified, there being no
certain representatives of either the ventral member, the presphenoid,
or the lateral members, the orbitosphenoids. On the dorsal side there
is, however, a large development of membrane bones. There is a large
_frontal_ (fig. 45, 3), unpaired, except in the embryo, united behind
with the parietal and postfrontal, and drawn out in front into a long
process which is overlapped by the prefrontals and posterior part of
the nasals. The frontal ends off freely below, owing to the
orbitosphenoidal region being unossified, it forms a considerable part
of the roof of the cranial cavity, but takes no part in the formation
of the wall.
Each _prefrontal_ (fig. 45, 14) forms part of the inner wall of the
orbit and sends ventralwards a process which meets the palatine.
The _postfrontals_ (fig. 44, 6) are small bones lying at the sides of
the posterior part of the frontal. Each is united with a number of
bones, on its inner side with the frontal and parietal, behind with
the squamosal, and ventrally with the alisphenoid. It also unites by
means of a strong descending process with an upgrowth from the jugal,
and thus forms a =postorbital bar= separating the orbit from the
lateral temporal fossa. The postfrontal forms also part of the outer
boundary of the supratemporal fossa.
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