The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
1. premaxillae (fused).
2. anterior nares.
3. nasal.
4. prefrontal.
5. frontal.
6. parietal.
7. maxillae.
8. transpalatine.
9. palatine.
10. pterygoid.
11. pro-otic.
12. exoccipital.
13. supra-occipital.
14. opisthotic.
15. epi-otic.
16. quadrate.
17. parasphenoid.
18. basisphenoid.
19. basi-occipital.
20. occipital condyle.
21. splenial.
22. dentary.
23. angular.
24. articular.
25. supra-angular.
26. coronoid.
27. vomer.
28. squamosal.
IX, X foramina for the ninth
and tenth cranial nerves.]
The disposition of the bones of the jaws is subject to much
modification in the Ophidia in order to adapt them for swallowing very
large prey. The arrangements again differ greatly in the venomous and
non-venomous snakes. In the non-venomous snakes, such as _Python_ and
_Tropidonotus_, the palatine is large and is fixed to the pterygoid
which extends outwards (fig. 51, 10) so as to be united to the
quadrate, and is at the same time firmly connected by the
transpalatine with the maxillae. The quadrate is united to the
squamosal, which is loosely attached to the cranium. The premaxillae is
moderately developed and bears teeth, and the maxillae forms a long bar
loosely connected with the rest of the skull. The rami of the mandible
are united only by an extremely elastic ligament. It is as regards the
maxillae and premaxillae that the skulls of venomous and non-venomous
snakes differ most. In the rattlesnake (_Crotalus_) and other venomous
snakes the premaxillae is extremely small and toothless. The maxillae is
small and subcylindrical, and is movably articulated to the lachrymal,
which also is capable of a certain amount of motion on the frontal.
The maxillae is connected by means of the transpalatine with the
pterygoid, which in its turn is united to the quadrate. When the mouth
is shut the quadrate is directed backwards, and carrying back the
pterygoid and transpalatine pulls at the maxillae and causes its
palatal face, to which the poison teeth are attached, to lie back
along the roof of the mouth. When the mouth opens the distal end of
the quadrate is thrust forward, and this necessitates the pushing
forward of the pterygoid and transpalatine, causing the tooth-bearing
surface of the maxillae to look downwards and the tooth to come into
the position for striking.
The Ophidian skull is also noticeable for the absence of the jugals
and quadratojugals. In poisonous snakes the place of the jugal is
taken by the zygomatic ligament which connects the quadrate and
maxillae.
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