The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
The pectoral girdle is well developed in all groups of reptiles except
the Ophidia, occurring even in the limbless Amphisbaenidae. It is very
solid in the Theromorpha. As a rule all three cartilage bones,
scapula, coracoid, and precoracoid are represented, and frequently
also the membrane bones,--clavicles, and interclavicle.
The coracoids are generally flat expanded bones, which sometimes, as
in Sauropterygia and Ichthyosauria, meet in a ventral symphysis;
sometimes, as in Lacertilia, are united with the sides of the sternum.
In Chelonia neither the coracoids nor precoracoids meet one another,
but their free ends are connected by fibrocartilaginous bands. In
Lacertilia the coracoids are pierced by fenestrae.
The precoracoid is generally represented, but the Theromorpha are the
only reptiles in which it is separately ossified; it forms a
well-marked process on the coracoid in Lacertilia (fig. 54, 5). It is
absent in Ichthyosauria, and Dinosauria, and probably in
Sauropterygia. In some Lacertilia and Chelonia the sternal ends of the
coracoids are unossified and form epicoracoids; in some Chelonia there
are also epiprecoracoids; but neither these nor the epicoracoids
overlap their fellows of the opposite side as they do in arciferous
Anura (see p. 185). In some Lacertilia with degenerate limbs the
pectoral girdle is also much reduced, in _Ophisaurus apus_ the ventral
borders of the coracoids are widely separated.
A scapula is always present, and is generally expanded distally, but
in the Chelonia the distal end is cylindrical. In the Theromorpha it
has an acromial process with which the precoracoid articulates, and it
is very large in Dinosauria. In the Chelonia the scapula and
precoracoid are ossified continuously. Among the Pterosauria,
_Pteranodon_ has an unique pectoral girdle; the scapula and coracoid
are ankylosed and the scapula articulates with the neural spines of
several ankylosed vertebrae.
Clavicles occur in some Theromorpha such as _Pariasaurus_, and also in
the Ichthyosauria, Sauropterygia, Rhynchocephalia, and most
Lacertilia. They are absent in the Pterosauria, the Chamaeleons among
Lacertilia, the Ophidia and the Crocodilia. They are wanting too in
the Chelonia, unless the first pair of ossifications in the plastron
are to be regarded as clavicles. In the Sauropterygia bones regarded
as the clavicles and interclavicle are generally well developed. The
unpaired ossification in the plastron of Chelonia is an interclavicle,
and a representative of the same bone occurs arising from the sternum
in Pterosauria. A well developed =T=-shaped interclavicle is found in
Ichthyosauria, Rhynchocephalia, Lacertilia, and some Theromorpha, such
as _Pariasaurus_.
THE LIMBS.
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