The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
This group of animals is very isolated, having no very close allies,
either living or extinct. The digits are provided with flat nails,
except the second digit of the pes, which is clawed. Canine teeth are
absent, and the dental formula is usually given as _i_ 1/2, _c_ 0/0,
_pm_ 4/4, _m_ 3/3. The upper incisors are long and curved, and have
persistent pulps as in Rodents; their terminations are, however,
pointed, not chisel-shaped, as in Rodents. The lower incisors have
pectinated edges. The grinding teeth have a pattern much like that in
_Rhinoceros_. In the skull (fig. 83) the postorbital processes of the
frontal and jugal almost or quite meet. The jugal forms part of the
glenoid cavity for articulation with the mandible, and also extends
forwards so as to meet the lachrymal. There is an alisphenoid canal.
There are as many as twenty-one or twenty-two thoracic vertebrae, and
the number of thoraco-lumbar vertebrae reaches twenty-eight or thirty.
There are no clavicles, and the scapula has no acromion; the coracoid
process is, however, well developed. The ulna is complete. In the
manus the second, third and fourth digits are approximately equal in
size, the fifth is smaller, and the first is vestigial. The femur has
a slight ridge representing the third trochanter. The fibula is
complete, but is generally fused with the tibia proximally. There is a
complicated articulation between the tibia and astragalus, which has a
pulley-like proximal surface. In the pes the three middle digits are
well developed, but there is no trace of a hallux, and the fifth digit
is represented only by a vestigial metatarsal.
The only representatives of the suborder are some small animals
belonging to the genus _Procavia_ (_Hyrax_), which is found in Africa
and Syria; some of the species are by many authors placed in a
distinct genus _Dendrohyrax_.
_Suborder_ (4). AMBLYPODA[130].
This suborder includes a number of primitive extinct Ungulates, many
of which are of great size. Their most distinguishing characteristics
are afforded by the extremities. In the carpus the bones interlock a
little more than is the case in most Subungulata, and the corner of
the os magnum reaches the scaphoid, while the lunar articulates
partially with both magnum and unciform, instead of only with the
magnum. In the tarsus the cuboid articulates with both the calcaneum
and the astragalus, which is remarkably flat. The manus and pes are
short, nearly or quite plantigrade, and have the full number of
digits. The cranial cavity is singularly small. Canine teeth are
present in both jaws, and the grinding teeth have short crowns, marked
by =V=-shaped ridges. The pelvis is large, the ilia are placed
vertically, and the ischia do not take part in the ventral symphysis.
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