The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
(_c_) The =horns of Rhinoceroses=. These are conical structures
composed of a solid mass of hardened epidermal cells growing from a
cluster of long dermal papillae. From each papilla there grows a fibre
which resembles a thick hair, and cementing the whole together are
cells which grow from the interspaces between the papillae. These
fibres differ from true hairs in not being developed in pits in the
dermis. Rhinoceros horns may be either one or two in number, and are
borne on the fronto-nasal region of the skull. They vary much in
length, the longest recorded having the enormous length of fifty-seven
inches.
(_d_) _Nails_, _hoofs_ and =claws=. In almost all mammals except the
Cetacea, these are found terminating the digits of both limbs. =Nails=
are more or less flattened structures, =claws= are pointed and
somewhat curved. In most mammals the nails tend to surround the ends
of the digits much more than they do in man. Sometimes the nail of one
digit differs from that of all the others; thus the second digit of
the pes in the Hyracoidea and Lemuroidea is terminated by a long claw,
the other digits having flat nails. In the Felidae the claws are
retractile, the ungual phalanx with claw attached folding back when
the animal is at rest into a sheath, above, or by the side of the
middle phalanx. In the Sloths and Bats enormously developed claws
occur, forming hooks by which the animals suspend themselves. In
_Notoryctes_ the third and fourth digits of the manus bear claws of
great size; similar claws occur in _Chrysochloris_, being correlated
in each case with fossorial habits. The nail at its maximum
development entirely surrounds the terminal phalanx of the digit to
which it is attached, and is then called a =hoof=. Hoofs are specially
characteristic of the Ungulata.
(_e_) =Spurs= and =beaks= are structures which are hardly represented
among mammals, while so characteristic of birds. They are however both
found in the Monotremata. In both _Echidna_ and _Ornithorhynchus_ the
male has a peculiar hollow horny spur borne on a sesamoid bone
articulated to the tibia. The jaws in _Ornithorhynchus_ are cased in
horny beaks similar to those of birds, and are provided with horny
pads which act as teeth.
(_f_) =Horny plates= of a ridged or roughened character occur upon the
anterior portion of the palate, and of the mandibular symphysis in all
three genera of recent Sirenia; also upon the toothless anterior
portion of the palate in Ruminants.
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