The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
Teeth are well developed in the vast majority of mammalia, and are of
the greatest morphological and systematic importance, many extinct
forms being known only by their teeth. Mammalian teeth differ from
those of lower animals in various well-marked respects. (1) They are
attached only to the maxillae, premaxillae and mandible, never to the
palatines, pterygoids or other bones. (2) They frequently have more
than one root. (3) They are always, except in some Odontoceti, placed
in distinct sockets. (4) They are hardly ever ankylosed to the bone.
(5) They are in most cases markedly heterodont. (6) They are commonly
developed in two sets, the milk dentition and permanent dentition.
It sometimes happens that teeth after being formed are reabsorbed
without ever cutting the gum. This is the case, for instance, with the
upper incisors of Ruminants.
The form of mammalian teeth varies much, some are simple conical
structures comparable to those of most reptiles, and these may either
have persistent pulps, as in the case of the upper canines of the
Walrus and the tusks of Elephants, or may be rooted as in most canine
teeth. Some teeth have chisel-shaped edges, and this may be their
original form, as in the human incisors, or may, as in those of
Rodents, be brought about by the more rapid wearing away of the
posterior edge, the anterior edge being hardened by a layer of enamel.
Then, again, the crown may, as in the majority of grinding teeth, be
more or less flattened. The various terms used in describing some of
the forms of the surface of grinding teeth are defined on page 345.
[Illustration FIG. 81. SKULL OF A YOUNG INDIAN RHINOCEROS (_R.
unicornis_), SHOWING THE CHANGE OF THE DENTITION × 1/7. (Brit. Mus.)
1. nasal.
2. frontal.
3. parietal.
4. zygomatic process of squamosal.
5. jugal.
_mI{1}._ milk incisor.
_mc._ milk canine.
_mpm{1}._ milk premolar.
_I{1}._ first incisor.
_c._ canine.
_pm{2}_, _pm{3}_, _pm{4}_. 2nd, 3rd and 4th premolars.
_m{1}_, _m{2}_. first and second molars.]
The teeth of the Aard Varks are compound, and differ completely from
those of all other mammals (see p. 425).
As a rule, the higher the general organisation of an animal the
better are its milk teeth developed, and the more do they form a
reproduction on a small scale of the permanent set. This fact is well
seen in the Primates, Carnivora and Ungulata. The method of notation
by which the dentition of any mammal can be briefly expressed as a
formula has been already described. The regular mammalian arrangement
of teeth for each side is expressed by the formula
_i_ 3/3 _c_ 1/1 _pm_ 4/4 _m_ 3/3 × 2; total, 44.
MONOTREMATA. In _Echidna_ teeth are quite absent. In the young
_Ornithorhynchus_[146] functional molar teeth of a multi-tubercular
type resembling those of some Mesozoic mammalia are present, but in
the adult they disappear, their office being discharged by horny
plates.
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