The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
_i_ 1/1 _c_ 0/0 _pm_ (0--1)/(0--1) _m_ 3/3, total 18 or 20.
The incisors always have chisel-like edges and persistent pulps, and
are separated by a wide diastema from the premolars. Canines are
always absent, and there are generally three grinding teeth not
preceded by milk teeth; their surface may be grooved, or may be
bunodont. Teeth are most numerous in the Duplicidentata (Hares and
Rabbits), in which the formula is _i_ 2/1 _c_ 0/0 _pm_ 3/2 _m_ 3/3,
total 28, and fewest in Hydromys and certain other forms, in which
the formula is _i_ 1/1 _c_ 0/0 _pm_ 0/0 _m_ 2/2, total 12. The hares
and rabbits are the only rodents which have well developed deciduous
incisors, though a vestigial milk incisor has been described in the
Mouse (_Mus musculus_). The last upper molar of _Hydrochaerus_ is very
complicated, its structure approaching that of the teeth of Elephants.
[Illustration FIG. 84. CARNASSIAL OR SECTORIAL TEETH OF CARNIVORA
(from FLOWER).
_Upper sectorial teeth_ of I. _Felis_, II. _Canis_, III. _Ursus_. 1.
anterior, 2. middle, 3. posterior cusp of blade, 4. inner lobe
supported on distinct root, 5. inner lobe posterior in position and
without distinct root, characteristic of the Ursidae.
_Lower sectorial teeth._ 1. _Felis_, 2. _Canis_, 3. _Herpestes_. 1.
anterior, 2. posterior lobe of blade, 3. inner tubercle, 4. heel.]
CARNIVORA have the teeth rooted and markedly diphyodont and
heterodont. The canines are greatly developed, and the incisors are
small.
In _CARNIVORA VERA_ the incisors are almost always 3/3. The fourth
upper premolar and first lower molar are differentiated as carnassial
teeth (see p. 436), and retain fundamentally the same characters
throughout the suborder. The upper carnassial (fig. 84, I. II. III.)
consists of a more or less compressed, commonly trilobed blade borne
on two roots, with an inner tubercle borne on a third root. The lower
carnassial has only two roots; its crown consists of a bilobed blade
with generally an inner cusp, and a heel or talon (fig. 84, 4) behind
the blade.
The most thoroughly carnivorous type of dentition is seen in the
Æluroidea, and especially in the cat tribe (Felidae). In the genus
_Felis_ the dental formula is _i_ 3/3 _c_ 1/1 _pm_ 3/2 _m_ 1/1, total
30. The incisors are very small, so as not to interfere with the
action of the large canines, the lower carnassial is reduced to simply
the bilobed blade (fig. 84, IV), and the cheek teeth are greatly
subordinated to the carnassial. The extinct _Machaerodus_ has the
upper canines comparable in size to those of the Walrus.
The Civets and Hyaenas have a dentition allying them closely to the
cats. The hyaena-like _Proteles_ has, however, the grinding teeth
greatly reduced.
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