The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
The mandible is rather short and broad, and the angle formed by the
meeting of the two rami is more obtuse than in most mammals. The
coronoid process is fairly well developed, and the angle is more or
less rounded. In most Primates the condyle is considerably widened,
but this is not the case in Man. In _Mycetes_ the mandible is very
large, its ascending portions being specially developed. The hyoid of
Primates is remarkable for the large expanded basi-hyal, which is
generally concave above and convex below. The anterior cornu is never
well ossified, but the thyro-hyal is always strong. In _Mycetes_ the
basi-hyal is enormously large, forming a somewhat globular thin-walled
capsule.
AUDITORY OSSICLES.
[Illustration FIG. 100. MALLEUS, STAPES AND INCUS OF _A._ MAN. _B._
DOG. _C._ RABBIT. (After DORAN) x 1.
1. head of malleus.
2. canal of stapes.
3. incus.
4. processus longus (or gracilis).
5. manubrium of malleus.
6. processus brevis.
7. lamella.]
There are in mammals four auditory ossicles forming a chain extending
from the fenestra ovalis to the tympanic membrane. Three of these, the
=malleus=, =incus= and =stapes=, are always distinct, while the
fourth, the =lenticular=, is smaller than the others and is sometimes
not distinct. The names are derived from human anatomy and indicate in
the case of the first three a more or less fanciful resemblance
respectively to a hammer, an anvil and a stirrup. The ossicles are
homologous as a whole to the hyomandibular of fishes and to the
columellar chain of Sauropsids and Amphibians. The malleus is
homologous to the extra-columella of Crocodiles and the stapes to the
columella. The =malleus= when typically developed consists of a
rounded _head_ (fig. 100, 1) which bears a surface articulating with
the incus, and a short _neck_ continued into a process, the
_manubrium_ (fig. 100, 5), which comes into relation with the tympanic
membrane. From the junction of the neck and manubrium two processes
are given off, a _processus longus_ or _gracilis_ (fig. 100, 4), which
in the embryo is continuous with Meckel's cartilage, and a _processus
brevis_ (fig. 100, 6). The =incus= generally consists of a more or
less anvil-shaped portion which articulates with the malleus, and of a
process which is connected with the stapes by the small =lenticular=.
The =stapes= is generally stirrup shaped, consisting of a basal
portion from which arise two _crura_ separated by a space the canal
through which a branch of the pharyngeal artery runs The lenticular is
frequently cartilaginous and sometimes is not developed at all.
The above is the arrangement of the auditory ossicles met with in the
higher Mammalia, but in the lower Mammalia the characters approach
more nearly to those met with in Sauropsids.
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