The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
By far the most primitive type of the pectoral or shoulder girdle is
found in the MONOTREMATA. The scapula (fig. 102, A, 1) is long and
recurved, and has only two surfaces, one corresponding to the
prescapular[1] fossa, the other to the postscapular[1] and
subscapular[169] fossae. The coracoid is a short bone attached above
to the scapula and below to the presternum; it forms a large part of
the glenoid cavity. In front of the coracoid there is a fairly large
flattened epicoracoid (fig. 102, 6); there is also a large =T=-shaped
interclavicle (fig. 102, 4), which is expanded behind and rests on the
presternum. The clavicles rest on and are firmly united to the
anterior border of the interclavicle. This shoulder girdle differs
greatly from that of any other mammals, and recalls that of some
Lacertilia.
[Illustration FIG. 102. _A_, SIDE VIEW, _B_, DORSAL VIEW OF THE
SHOULDER GIRDLE AND PART OR THE STERNUM OF THE SPINY ANTEATER
(_Echidna aculeata_) × 1. (After PARKER.)
1. scapula.
2. suprascapula.
3. clavicle.
4. interclavicle.
5. coracoid.
6. epicoracoid.
7. glenoid cavity.
8. presternum.
9. second sternal rib.
10. second vertebral rib.]
In MARSUPIALS, as in all mammals except the Monotremes, the shoulder
girdle is much reduced; there are no epicoracoids and interclavicle,
and the coracoid forms simply a small process on the scapula,
ossifying from a centre separate from that giving rise to the rest of
the bone. The scapula has a long acromion, and a clavicle is always
present except in _Perameles_. Unossified remains of the precoracoids
are found at either end of the clavicle. The scapula of _Notoryctes_
has a very high overhanging spine, and there is a second strong ridge
running along the proximal part of the glenoid border.
The shoulder girdle of the EDENTATA shows some very curious
variations. In _Orycteropus_ the scapula is of very normal form and
the clavicle is well developed. In the Pangolins and Anteaters the
scapula is very broad and rounded; there is no clavicle in the
Pangolins, and generally only a vestigial one in Anteaters. In
Armadillos, Sloths, and Megatheriidae, the acromion is very long and
the clavicle is well developed. In the Sloths, _Megatherium_, and
_Myrmecophaga_, a connection is formed between the coracoid, which is
unusually large, and the coracoid border of the scapula, converting
the coraco-scapula notch into a foramen. In _Bradypus_ the clavicle is
very small, and is attached to the coracoid, which sometimes forms a
distinct bone[170].
In the SIRENIA the scapula is somewhat narrow and curved backwards:
the spine, acromion, and coracoid process are moderately developed,
and there is no clavicle.
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