The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
MONOTREMATA. In _Echidna_ the carpus is broad, the scaphoid and lunar
are united and there is no centrale. The pisiform is large and several
other sesamoid bones occur. Each of the five digits is terminated by a
large ungual phalanx. In _Ornithorhynchus_ the manus is more slender,
but the general arrangement is the same as in _Echidna_.
MARSUPIALIA. The carpus has no centrale and the lunar is generally
small or absent. Five digits are almost always present. In _Choeropus_
however the only two functional digits are the second and third, which
have very long closely apposed metacarpals; the fourth digit is
vestigial, but has the normal number of phalanges, while the first and
fifth are absent. The manus in _Notoryctes_ is extraordinarily
modified, the scaphoid and all the distal carpalia are apparently
fused, the first, second, and fifth digits are very small, the third
and fourth, though having only one phalanx apiece, bear each an
enormous claw. Lying on and obscuring the ventral surface of the manus
is a large bone, probably a sesamoid.
Among the EDENTATA there is a great diversity in the structure of the
manus, the centrale is however always wanting, and except in _Manis_
the scaphoid and lunar are distinct. In the Sloths the manus is very
long, narrow, and curved, and terminated by two or three long hooked
claws, borne by the second and third, or the second, third and fourth
digits. The fifth digit is absent, and the fourth is represented only
by a small metacarpal. In the Anteaters the third digit is greatly
developed and bears a long hooked claw. In _Myrmecophaga_ all five
digits are fairly well though irregularly developed, in _Cycloturus_
the first, fourth, and fifth, are vestigial. In the Armadillos the
manus is broad, and has strongly developed ungual phalanges. The
digits, though almost always five in number, vary much in their
relative arrangement. In _Dasypus_ they are regular, but are
remarkably irregular in Priodon. The pollex is absent in Glyptodonts
and in _Megatherium._ In _Megatherium_ the fifth digit is clawless
while the second, third, and fourth bear enormous claws. In the
Manidae the scaphoid and lunar are united; five digits are present,
the third and fourth being very large, and all being terminated by
deeply cleft ungual phalanges. In _Orycteropus_ the pollex is absent,
while the other digits are terminated by pointed ungual phalanges.
In SIRENIA the general structure of the manus is quite of the ordinary
mammalian type. In _Manatus_ most of the bones of the carpus are
distinct, but in _Halicore_ many, especially those of the distal row,
have coalesced. The digits are always five in number and have the
normal number of flattened phalanges.
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