The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
1. head.
2. great trochanter.
3. lesser trochanter.
4. third trochanter.
5. shaft.
6. condyles.]
In the Mystacoceti among the CETACEA small nodules of bone or
cartilage occur connected with the vestigial pelvis, and may represent
the femur and tibia. No trace of the skeleton of the hind limb is
known in the Odontoceti.
In the UNGULATA VERA the femur is noticeable for the size of the great
trochanter (fig. 109, 2); there is no definitely constricted neck
separating the head from the rest of the bone, and the lesser
trochanter (fig. 109, 3) is not very prominent. All Perissodactyles
except the Chalicotheriidae show a strongly marked third trochanter,
but this is absent in all known Artiodactyles. The development of the
fibula in general corresponds to that of the ulna. In _Rhinoceros_,
_Macrauchenia_, _Tapirus_ and the Suina it is distinct and fairly well
developed; in the Tragulina on the other hand it is vestigial, being
reduced to the proximal end only. In the Ruminantia and Tylopoda also,
it is much reduced forming merely a small bone attached to the distal
end of the tibia, sometimes, as in the Red deer a slender vestige of
the proximal end also is preserved quite detached from the distal
portion; in the Horse this proximal portion is all that there is found
of the fibula. The progressive diminution of the fibula can be well
seen in the series of forms that are regarded as the ancestors of the
Horse. The patella of the Ungulata vera is well ossified, but
fabellae[177] are not usually found.
SUBUNGULATA. Of the Toxodontia, _Toxodon_ has no third trochanter
while _Typotherium_ and _Astrapotherium_ have one. In the Condylarthra
the femur has well-marked lesser and third trochanters, and the fibula
and patella are well developed. In the Hyracoidea there is a slight
ridge on the femur in the place of the third trochanter, the fibula is
complete, but is generally fused to the tibia at its proximal end.
Of the Amblypoda, _Coryphodon_ has a third trochanter, but
_Uintatherium_ has none; in this respect, in the vertical position and
general appearance (fig. 108) of the limb, and in the articulation of
the fibula with the calcaneum, the leg of _Uintatherium_ closely
approaches that of the Proboscidea.
In the Proboscidea the femur is very long and straight, the
development of trochanters is slight, and the fibula though slender is
complete and articulates with the calcaneum.
A third trochanter is found in the Tillodontia.
In RODENTIA the femur is variable, the great trochanter is generally
large and so sometimes is the third as in the Hares. In most Rodents
as in the Beaver the fibula is distinct, sometimes as in the Hares it
is united distally with the tibia. The patella is well developed, and
so too are the fabellae as a general rule.
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