The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
A. _Perennibranchiata_, whose chief distinguishing skeletal characters
are that the skull is elongated, the premaxillae are not ankylosed, the
maxillae are vestigial or absent; there are sometimes no nasals, and
the palatines bear teeth;
e.g. _Siren_, _Proteus_, _Menobranchus_.
B. _Derotremata_, whose chief distinguishing skeletal characters are
that there are large maxillae and nasals; teeth are borne by both
maxillae and premaxillae; there are no palatines; and both pectoral and
pelvic limbs are always present;
e.g. _Amphiuma_, _Megalobatrachus_, _Cryptobranchus_.
_Suborder_ (2). SALAMANDRINA.
The vertebrae are opisthocoelous. The skull is broad, and teeth are
borne by both premaxillae and dentaries. Nasal bones are present. The
remains of only two branchial arches are found in the adult. The
carpus and tarsus are more or less ossified.
This suborder includes the Newts (_Molge_), Salamanders
(_Salamandra_), and _Amblystoma_.
_Order_ 2. LABYRINTHODONTIA[52].
These are extinct Amphibia with a greatly developed dermal
exoskeleton, which is generally limited to the ventral surface. The
body and tail are long and in some cases limbs are absent. The teeth
are pointed and often have the dentine remarkably folded. The
vertebrae are amphicoelous, and are generally well ossified. The skull
is very solid, and has a greatly-developed secondary roof which hides
the true cranium and is very little broken up by fossae. Paired
dermal supra-occipitals are found, and there is an interparietal
foramen. The epi-otics and opisthotics form a pair of bones distinct
from the exoccipitals. Four simple limbs of moderate length are
generally present, and in some cases all four limbs are
pentedactylate. Among the better known genera of Labyrinthodonts are
_Mastodonsaurus_, _Nyrania_, and _Archegosaurus_.
_Order 3._ GYMNOPHIONA[53].
These animals form a group of abnormal worm-like Amphibia having an
exoskeleton in the form of subcutaneous scales arranged in rings. The
vertebrae are biconcave and are very numerous; very few however belong
to the tail. The skull has a complete secondary bony roof, the
mandible bears teeth and has an enormous backward projection of the
angular. The hyoid arch has very slender cornua and no distinct body,
it is attached neither to the cranium nor to the suspensorium. The
ribs are very long and there are no limbs or limb girdles.
_Order 4._ ANURA.
These are tailless Amphibia, which except in a few instances, are
devoid of an exoskeleton. The vertebrae are as a rule procoelous, and
are very few in number. The post-sacral part of the spinal column
ossifies continuously, forming an unsegmented cylindrical rod, the
urostyle. Remains of the notochord persist, lying _vertebrally_, i.e.
enclosed within the centra of the several vertebrae, and not as in
Urodela lying between one vertebra and the next. The skull is very
short and wide. The mandible is almost always, if not invariably,
toothless.
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