The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
The =tarsus= consists of eight small bones arranged in a proximal row
of three, the =tibiale=, =intermedium= and =fibulare=, and a distal
row of four =tarsalia=, with one bone, the =centrale= (fig. 23, A, 7),
interposed between the two rows. In the specimen examined, the
=tibiale=, is a small bone articulating with the tibia, the
=intermedium= (fig. 23, A, 5) is larger and articulates with both
tibia and fibula, the =fibulare= is the largest of the three and
articulates with the fibula.
The bones of the distal row are =tarsalia 1=, =2=, =3=, and a bone
representing =4= and =5= fused. In the specimen examined tarsale 1 is
pushed away dorsally (fig. 23, A, 8), so as to lie between the tibiale
and tarsale 2. All the tarsal bones are small and somewhat polygonal,
and are connected with one another, and with the tibia and fibula on
the one hand, and with the metatarsals on the other by a thin layer of
cartilage.
The five =digits= of the foot each consist of a =metatarsal= and of a
certain number of =phalanges=. In the specimen examined, owing to the
shifting of tarsale 1, the first metatarsal as well as the second
articulates with tarsale 2, while the fifth metatarsal articulates
partially with the bone representing the fused tarsalia 4 and 5,
partially with the fibulare. All the bones of the digits except the
distal phalanges are terminated at each end by cartilaginous
epiphyses, the distal phalanx of each digit has a cartilaginous
epiphysis only on its proximal end.
The first, second, and fifth digits have two phalanges apiece, the
third and fourth have three.
Figure 31 B, showing a Newt's tarsus copied from Gegenbaur, has
precisely the arrangement generally regarded as primitive for the
higher vertebrates, except that tarsalia 4 and 5 are fused.
FOOTNOTES:
[54] i.e. between one vertebra and the next.
CHAPTER XI.
THE SKELETON OF THE FROG[55] (_Rana temporaria_).
I. EXOSKELETON.
The skin of the frog is smooth and quite devoid of scales or other
exoskeletal structures. The only exoskeletal structures met with in
the frog are:--
1. The =teeth=, which are most conveniently described with the
endoskeleton.
2. The horny covering of the calcar or prehallux (see p. 167).
II. ENDOSKELETON.
The endoskeleton of the adult frog consists partly of cartilage,
partly of bone and each of these types of tissue occurs in two forms.
The cartilage may be hyaline, as in the omosternum and xiphisternum,
or may be more or less calcified as in part of the suprascapula and
the epiphyses of the limb bones. The bone may be cartilage bone, or
membrane bone.
The skeleton is divisible into an =axial portion= consisting of the
skull, vertebral column, and sternum, and an =appendicular portion=
consisting of the skeleton of the limbs and their girdles.
1. THE AXIAL SKELETON.
A. THE VERTEBRAL COLUMN.
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