The Wonders of Life: A Popular Study of Biological PhilosophyHaeckel, Ernst
Philosophy
The Wonders of Life: A Popular Study of Biological Philosophy
Haeckel, Ernst
Biology; Evolution (Biology); Life -- Origin
In the higher animals and plants we usually regard as the "real
individual" (in the wider sense of the word) the tissue-forming
organism made up of various organs; and we may here briefly and
instructively call this the histonal individual (or, more briefly,
the "histonal"). Botanists call this individual phenomenon among
the metaphyta a sprout (_blastus_). Zoologists give the title of
"person" (_prosopon_) to the corresponding unity among the animals.
The two forms agree very much in their general features, and may be
called "individuals of the second order," if we take the cells to be
the first and the stock the third stage in the hierarchy of organic
individuality. In comprising them here under the general head of
histonals, or histonal individuals, I mean by this to designate the
definite physiological unity of the multicellular and tissue-forming
organism, as contrasted with the unicellular protist on the one hand,
and the higher stem, made up of several histonals, on the other.
The plant-histonal, which Alexander Braun especially clearly marked
out and described as the sprout, is found in two principal forms in
the kingdom of the metaphyta--the lower form of the layer-sprout
(_thallus_) and the higher form of the stalk-sprout (_culmus_).
The thallus predominates in the lower and older sub-kingdom of the
layer-plants (_thallophyta_), in the classes of the algæ and fungi;
the culmus in the higher and younger sub-kingdom of the stalk-plants
(_cormophyta_), in the classes of the mosses, ferns, and flowering
plants. The culmus presents in general the characteristic form of
an axial central organ, the stalk, with lateral organs, the leaves,
attached to this at the sides, the former having an unlimited vertical
growth and the latter an unlimited basal growth. The thallus does not
yet show this important morphological division. There are, however,
exceptions in both groups of the metaphyta. The large and highly
developed _fucoidea_ among the algæ exhibit similar differentiations
of organs to those we distinguish as stalk and leaves in the higher
cormophyta. On the other hand, they are wanting in the lower
liverworts, which form a thallus like many of the algæ; thus, for
instance, the liverwort _riccia fluitans_ is just like the brown
alga _dictyota dichotoma_. Other primitive liverworts (such as the
_anthoceros_) have also a very simple thallus; but most of them have
a separation of the thallus into an axial organ (stalk) and several
lateral organs (leaves). In the distribution of labor among the
leaves there then emerge the differences between the lower leaves,
foliage leaves, higher leaves, and flower leaves. A simple poppy-plant
(_papaver_) or a single-flowered _gentiana ciliata_, which has only one
bloom at the top of its branchless stalk, is a good example of a highly
developed culmus.
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