The Wonders of Life: A Popular Study of Biological PhilosophyHaeckel, Ernst
Philosophy
The Wonders of Life: A Popular Study of Biological Philosophy
Haeckel, Ernst
Biology; Evolution (Biology); Life -- Origin
III. CENTROPLANE TYPES.--The natural middle of the body is a plane,
the median or chief plane (_planum medianum_ or _sagittale_); it
divides the bilateral body into two symmetrical halves, the right
and the left. With this is associated the characteristic antithesis
of back (_dorsum_) and belly (_venter_); hence, in botany this
type (found, for instance, in most green leaves) is called the
dorsiventral, and in zoology the bilateral in the narrower sense. One
characteristic of this important and wide-spread type is the relation
of three different axes, vertical to each other; of these three
straight axes (enthyni) two are unequipolar and the third equipolar.
Hence, the centroplanes may also be called tri-axial (_triaxonia_).
In most of the higher animals (as in our own frame) the longest of
the three axes is the principal one (_axon principalis_); its fore
pole is the oral or mouth pole, and its hinder pole is the aboral
or caudal (tail) pole. The shortest of the three enthyni is, in our
body, the sagittal (arrow) or dorsiventral axis; its upper pole is
at the back and its lower pole at the belly. The third axis--the
transverse or lateral axis--is equipolar, one pole being called the
right and the other the left. The various parts which make up the two
halves of the body have relatively the same disposition in each half;
but absolutely speaking (namely, in relation to the middle plane)
they are oppositely arranged.
Further, the centroplane or bilateral forms are also characterized
by three vertical axes which may be drawn through each of the normal
axes. The first of these normal planes is the median plane; it is
defined by the chief axis and the sagittal axis, and divides the body
into two symmetrical halves, the right and left. The second normal
plane is the frontal plane; this passes through the chief axis and
the transverse axis (which is parallel to the frontal surface in
our body), and divides the dorsal half from the ventral half. The
third normal plane is the cingular (waist) plane: this is defined by
the sagittal and transverse axes. It divides the head half (or the
vertical part) from the tail half (or the basal part).
The name "bilateral symmetry," which is especially applied to the
centroplane and dorsiventral types, is ambiguous, as I pointed out
in 1866 in an exhaustive analysis and criticism of these fundamental
forms in the fourth book of the _General Morphology_. It is used in
five different senses. For our present general purpose it suffices to
distinguish two orders of centroplane types, the bilateral-radial and
the bilateral-symmetrical; in the former the radial (pyramidal) form
is combined with the bilateral, but not in the latter.
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