The Wonders of Life: A Popular Study of Biological PhilosophyHaeckel, Ernst
Philosophy
The Wonders of Life: A Popular Study of Biological Philosophy
Haeckel, Ernst
Biology; Evolution (Biology); Life -- Origin
The classes of the lower animals which we comprise under the name
cœlenteria (or cœlenterata in the widest sense) generally agree
in having all the functions of nutrition accomplished exclusively (or
for the most part) by a single system of organs, the gastro-canal or
gastro-vascular system. From their common stem-group, the gastræads,
three different stems have been evolved--the sponges, cnidaria, and
platodes. All these cœlenteria have three features in common: (1)
The gastric canal or tube has only one opening--the primitive mouth,
which serves at once for admitting food and ejecting indigestible
matter; there is no anus; (2) there is no special body-cavity
(_cœloma_) distinct from the gastric tube; (3) there is also no
trace of a vascular system. All cavities that are found in these lower
animals besides the digestive gut-cavity are direct processes from it
(with the exception of the nephridia in the platodes).
While the simple digestive gut is the sole organ of nutrition in the
stem-group of the gastræads, we find other structures co-operating in
the rest of the cœlenteria. The characteristic stem of the sponges
is distinguished by the piercing of the wall of the gastric vesicle
with several holes. Through these water pours into the body, bringing
with it the small particles of food which are received and digested
by the ciliated cells of the entoderm; the water emerges again by the
mouth-opening (_osculum_). The best-known of the sponges is the common
bath-sponge (_euspongia officinalis_), the horny skeleton of which
we use daily in washing. In these and most other sponges the large,
unshapely body is traversed by a number of branching canals, on which
there are thousands of tiny vesicles, produced by the multiplication
of a simple gastric vesicle of the primitive sponge (_olynthus_). Each
of these ciliated chambers is really a tiny gastræa, a "person" of the
simplest character (_cf._ chapter vii.). Hence we may regard the whole
sponge-body as a gastræad-stock (_cormus_).
The large group of the cnidaria offers a long series of evolutionary
stages, from very small and simple to very large and elaborate forms.
Some of them remain at a very low stage, as does our common green
fresh-water polyp (_hydra viridis_), which only differs from the
gastræa by a few variations in tissue and the formation of a crown of
feelers about the mouth. Most of the polyps form stocks (_cormi_),
the individuals shooting out buds which remain joined to the mother
animal. In these and all the other stock-forming animals the nutrition
is communistic; all the food that the individuals get and digest is
conducted by tubes to the common fund and equally distributed. In all
the larger cnidaria the body-wall becomes thicker, and is traversed by
branching gastro-canals; these convey the nutritive fluid to all parts
of the body.
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