The Wonders of Life: A Popular Study of Biological PhilosophyHaeckel, Ernst
Philosophy
The Wonders of Life: A Popular Study of Biological Philosophy
Haeckel, Ernst
Biology; Evolution (Biology); Life -- Origin
Gastric or internal lungs are only found in the higher animals, to
which we give the name of quadrupeds (or _tetrapoda_), the amphibia and
amniotes, and their fishlike ancestors, the dipneusta. These internal
lungs are sac-shaped folds of the fore-gut, formed originally from the
swimming-bladder (_nectocystis_) of the fishes by change of function.
This air-filled bladder, a sac-shaped appendage of the gullet, merely
serves the purpose of a hydrostatic organ, by varying the specific
weight, in the fishes. When the fish wishes to descend it contracts
the bladder and becomes heavier; it rises to the top by inflating it
again. The lungs were formed by the adaptation of the blood-vessels
in the wall of the swimming-bladder to the interchange of gases. In
the oldest living lung-fishes (_ceratodus_) it is still a simple sac
(_monopneumones_=one-lunged); in the others the simple gullet-cavity
divides early into a pair of sacs (_dipneumones_, two-lunged). The
wind-pipe (_trachea_--not to be confused with the organ of the same
name in the tracheata) is formed by the lengthening of their stalk
and strengthening of it with cartilaginous rings. At the anterior end
of the trachea we find already formed in the amphibia the larynx, the
important organ of voice and speech.
The function of removing unusable matter is not less important to the
organism than breathing. Just as breathing gets rid of the poisonous
carbonic acid, so the kidneys remove fluid and solid excreta in the
shape of urine; these are partly acid (uric acid, hippuric acid, etc.),
partly alkaline (urea, guanine, etc.). In most of the cœlomaria
special organs for removing these would be superfluous, as this is
accomplished (like breathing) by the stream of water that is constantly
passing through the whole body. But with the platodes we begin to
find important excretory organs in the nephridia, a pair of simple
and ramified canals which lie on either side of the gut, and open
outward. These primitive renal canals are transmitted by the platodes
to the vermalia, and by these to the higher stems of the cœlomaria.
In the latter they generally open by special funnels into the inner
body-cavity, which serves as first receptacle for the urine. Their
outer opening sometimes (primarily) goes through the outer skin at the
back (excretory pores), sometimes (secondarily) to the rectum, and
so out through the anus. The oldest articulates, the annelids, have
a pair of nephridia in each segment of the body; each renal canal,
or segmental canal, consists of three sections, an inner funnel
which opens into the body-cavity, a middle glandular section, and an
external bladder that ejects the urine by contraction. The disposition
of the renal system in the internally articulated vertebrates is very
similar to this; but now complicated structures begin to appear, a
pair of compact kidneys (_renes_), which are made up of a number of
branching nephridia. Three generations of kidneys succeed each other,
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