The Wonders of Life: A Popular Study of Biological PhilosophyHaeckel, Ernst
Philosophy
The Wonders of Life: A Popular Study of Biological Philosophy
Haeckel, Ernst
Biology; Evolution (Biology); Life -- Origin
The class of crustacea affords the most numerous and most instructive
examples of degeneration through parasitism, because in this class it
is found polyphyletically in very different orders and families, and
because their highly organized body shows every stage of degeneration
together in the different organs. The free-living crustacea generally
move about very rapidly and ingeniously; their numerous bones are
well jointed and excellently adapted for the most varied methods of
locomotion (running, swimming, climbing, digging, etc.); their organs
of sense are highly developed. As these are no longer used when
they take to parasitism, they atrophy and gradually disappear. The
younger crustacea all proceed from the same characteristic form of
the _nauplius_, and swim freely about; later, when they settle down
to parasitic habits, their organs of sense and locomotion atrophy. As
Fritz Müller-Desterro showed in his famous little work, _For Darwin_
(1864), forty years ago, the crustacea afford most luminous proofs of
the theory of descent and selection, and of progressive heredity and
the biogenetic law. These facts are the more important as the crab
undergoes the same degeneration by parasitic habits in a number of
different orders and families.
From parasitism we must entirely distinguish that intimate life-union
of two different organisms which we called symbiosis or mutualism.
Here we have an association of two living things for their mutual
benefit, while the parasite lives entirely at the expense of his
host. Symbiosis is found among the protista, being very wide-spread
among the radiolaria. In the gelatinous envelope (_calymma_) which
encloses the central capsule of their unicellular bodies we find a
number of motionless yellow cells (_zooxanthella_) scattered. These
are protophyta or (as it is said) "unicellular algæ" of the class of
paulotomea (_palmellacea_). They receive protection and a home from the
radiolaria, grow plasmodomously, and multiply by rapid segmentation.
A large part of the starch-flour and the plasm which they form by
carbon-assimilation goes as food directly to the radiolarium-host; the
other part of the xanthella goes on growing and multiplying. Similar
yellow zooxanthella or green zoochlorella are found as symbionta in the
tissues of many animals. Our common fresh-water polyp (_hydra viridis_)
owes its green color to the zoochlorella which live in great numbers
on the ciliated cells of its entoderm (the digestive gut-epithelium).
In general, however symbiosis is rarer in the metazoa than in the
metaphyta. In the latter case it is the fundamental feature of a whole
class of plants, the lichens. Each lichen consists of a plasmodomous
plant (sometimes a protophyte, sometimes an alga) and a plasmophagous
fungus. The latter affords a home, protection, and water to the green
alga, which repays the service by providing food.
XI
REPRODUCTION
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