The Wonders of Life: A Popular Study of Biological PhilosophyHaeckel, Ernst
Philosophy
The Wonders of Life: A Popular Study of Biological Philosophy
Haeckel, Ernst
Biology; Evolution (Biology); Life -- Origin
Asexual or monogenetic tocogony (also called "vegetative
multiplication") is always effected by a single organic individual,
and so must be traced to its transgressive growth. When this affects
the entire body as a total growth, the whole dividing into two or
more equal parts, we call the monogenetic process division (or
segmentation). But when the growth is partial, and affects only a
part of the individual, or when this special part separates from
the generating organism in the form of a bud (_gemma_), the process
is called budding or gemmation (_gemmatio_). Hence the essential
difference between the two forms of generation is that in division the
parent disappears in its partial products (children); these are of
the same age and form. But in budding the generating parent retains
its individuality; it is larger and older than the young bud. This
important difference between division and gemmation, which is often
overlooked, holds good both for protists (unicellulars) and histona
(multicellulars). The fact that in division the individual as such
is destroyed is a sufficient refutation of Weismann's theory of the
immortality of the unicellulars. (See above, and also the _Riddle_,
chapter xi.)
Reproduction by division is by far the most common of all forms of
propagation. It is the normal form of monogony, not only in many of the
protists, but also in the tissue-cells which compose the tissues of
the histona. It is, moreover, the sole method of propagation for most
of the monera, both chromacea and bacteria, which are in consequence
often comprised under the title of "cleavage-plants" (_schizophyta_).
Self-cleavage is also found among the higher multicellular
organisms--namely, the cnidaria (polyps, medusæ). It usually takes the
form of division into two parts (_dimidiatio_ or hemitomy), the body
splitting into two equal halves. The plane of division is sometimes
indefinite (fragmentary-cleavage), sometimes coincident with the long
axis (length-cleavage), sometimes with the transverse axis, vertical
to the long axis (transverse-cleavage), and less frequently with a
diagonal axis (oblique-cleavage). When the segmentation of a cell
proceeds so rapidly that the transverse-cleavage follows immediately
on the length-cleavage, and the two are at length made to coincide,
twofold division is changed into fourfold division. And when the
process is repeated in quick succession, and the body falls at last
into a number of small and equal pieces, we have manifold division
(polytomy); as in the spore-formation of the sporozoa and rhizopoda,
and in the embryonic sac of the phanerogams.
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