The Wonders of Life: A Popular Study of Biological PhilosophyHaeckel, Ernst
Philosophy
The Wonders of Life: A Popular Study of Biological Philosophy
Haeckel, Ernst
Biology; Evolution (Biology); Life -- Origin
In a large number of lower animals and plants sexual and asexual
generation regularly alternate. Among the protists we find this
alternation of generation in the sporozoa; among the metaphyta in the
mosses and ferns; and among the metazoa in the cnidaria, platodes,
tunicates, etc. Often the two generations differ considerably in shape
and degree of organization. Thus, in the mosses the asexual generation
is the spore-forming moss capsule (_sporogonium_), while the sexual is
the moss plant with stalk and leaves (_culmus_). In the case of the
ferns, on the other hand, the latter is spore-forming and monogenetic,
while the thallus-formed, simple, and small fore-germ (_prothallium_)
is sexually differentiated. In most of the cnidaria a small stationary
polyp is developed out of the ovum of the free-swimming medusa, and
this polyp, in turn, generates by budding medusæ, which reach sexual
maturity. In the tunicates (salpa) a sexual social form alternates with
an asexual solitary form; the chain-salpa of the former are smaller and
differently shaped than the large individual salpa of the latter, which
again generate chains by budding. This special form of metagenesis was
the first to be observed, as it was in 1819 by the poet Chamisso, when
he sailed round the world. In other cases (for instance, in the closely
related _doliolum_) a sexual generation alternates with two (or more)
neutral ones. The explanation of these various forms of alternating
generations is given in the laws of latent heredity (atavism), division
of labor, and metamorphosis, and especially by the biogenetic law.
While in real metagenesis (alternation of generations in the strict
sense) the asexual generation propagates by budding or spore-formation,
this is done parthenogenetically in the cognate process of
heterogenesis. This it is which, especially in many of the articulates,
causes an immense increase of the species in a short time. Among the
insects we have the leaf-lice (aphides), and among the crustacea the
water-fleas (daphnides), that propagate in great numbers during warm
weather by unfertilized "summer-ova." It is not until the autumn that
males appear and fertilize the large "winter-ova"; in the following
spring the first parthenogenetic generation issues from the winter
eggs. The two heterogenetic generations are very different in the
parasitic suctorial worms (trematodes). From the fertilized ovum of the
hermaphrodite distoma we get simply constructed nurses (pædogenetic
larvæ), inside which cercaria are generated from unfertilized ova;
these travel, and are afterwards converted (inside another animal) into
distoma once more.
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