The Wonders of Life: A Popular Study of Biological PhilosophyHaeckel, Ernst
Philosophy
The Wonders of Life: A Popular Study of Biological Philosophy
Haeckel, Ernst
Biology; Evolution (Biology); Life -- Origin
The remarkable stem of the sea-dwelling echinoderms or "prickly skins"
is distinguished from all the other animal groups by a number of
striking peculiarities; prominent among these are the special formation
of their active and passive motor organs and the curious form of their
individual development. In this ontogenesis two totally different
forms appear successively--the simple astrolarva and the elaborately
organized and sexually mature astrozoon. The small, free-swimming
astrolarva has the general structural features of the rotatoria, and
so shows, in accordance with the biogenetic law, that the original
stem-form of the echinoderms (the amphoridea) belonged to this group
of the vermalia. I have briefly explained these structures in the
_History of Creation_ (chapter xxii.), and more fully in my essay on
the amphoridea and cystoidea (1896). The little astrolarva has no
muscles, and no water-vessels or blood-vessels. It moves by means
of vibratory lashes or bands, which are attached to special armlike
processes at the surface. These arms are regularly developed to the
right and left of the bilateral symmetrical larva (which as yet shows
no trace of the five-rayed structure). By a very curious modification
the small bilateral astrolarva is transformed into the totally
different pentaradial astrozoon, the large sexually mature echinoderm
with a pronounced five-rayed structure. (See _Art-forms in Nature_,
plates 10, 20, 30, 40, 60, 70, 80, 90, and 95.) It has a most elaborate
organization, with muscles and cuticular skeleton, blood-vessels and
water-vessels, etc. A section of the astrozoa--the living crinoidea,
or sea-lilies, and the extinct classes of blastoidea (sea-buds),
cystoidea (sea-apples), and amphoridea (sea-urns)--grow in stationary
fashion at the bottom of the sea. The other four extant classes
creep about in the sea--the sea-gherkins (holothuria), the star-fish
(asteridea and ophoidea), and the sea-urchins (echinidea). Their
creeping motion is accomplished by two kinds of organs--water-feet
and skin-muscles. The latter find their support and attachment in
solid calcareous needles, which develop from chalky deposits in the
corium. As these calcareous needles (which are particularly conspicuous
in the sea-urchin) are set movably in special protuberances of the
calcareous plates of the cuticular skeleton, and moved by little
muscular needles, the echinoderms walk on them as if they were stilts.
Between these, however, a number of water-feet arise from inside--thin
tubes like the fingers of a glove, which are filled with water by
an internal conduit-system (the so-called ambulacral system) and
become stiff. These very extensible ambulacral feet, often provided
with a suctorial plate at the closed outer end, serve for creeping,
sucking, touching, and grasping. As these distinctive motor organs
of the echinoderms--both the ambulacral feet with their complicated
water-tubes and the movable needles with their joints and muscles--are
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