This fact, especially when taken in connection with the formation of fresh segments outside the germinal disc and with other facts which I shall mention hereafter, is of great morphological interest as bearing upon the nature and homologies of the food-yolk. It also throws light upon the behaviour and mode of increase of the nuclei. All the nuclei, both those of the segments and those of the yolk, have the peculiar structure I described in the last stage. In specimens of this stage I have been able to observe certain points which have an important bearing upon the behaviour of the nucleus during cell-division. Three figures, illustrating the behaviour of the nucleus, as I have seen it in sections of blastoderms hardened in chromic acid, are shewn in Pl. 6, figs. 7_a_, 7_b_ and 7_c_. In the place of the nucleus is to be seen a sharply defined figure (Fig. 7_a_) stained in the same way as the nucleus or more deeply. It has the shape of two cones placed base to base. From the apex of each cone there diverge towards the base a series of excessively fine striæ. At the junction between the two cones is an irregular linear series of small deeply stained granules which form an apparent break between the two. The line of this break is continued very indistinctly beyond the edge of the figure on each side. From the apex of each cone there diverge outwards into the protoplasm of the cell a series of indistinct markings. They are rendered obscure by the presence of yolk-spherules, which completely surround the body just described, but which are not arranged with any reference to these markings. These latter striæ, diverging from the apex of the cone, are more distinctly seen when the apex points to the observer (Fig. 7_b_), than when a side of the cone is in view. The striæ diverging outwards from the apices of the cones must be carefully distinguished from the striæ of the cones themselves. The cones are bodies quite as distinctly differentiated from the protoplasm of the cell as nuclei, while the striæ which diverge from their apices are merely structures in the general protoplasm of the cell. In some cells, which contain these bodies, no trace of a commencing line of division is visible. In other cases (Fig. 7_c_), such a line of division does appear and passes through the junction of the two cones. In one case of this kind I fancied I could see (and have represented) a coloured circular body in each cone. I do not feel any confidence that these two bodies are constantly present; and even where visible they are very indistinct. Instead of an ordinary nucleus a very indistinctly marked vesicular body sometimes appears in a segment; but whether it is to be looked on as a nucleus not satisfactorily stained, or as a nucleus in the act of being formed, I cannot decide.