Calamites are among the most abundant fossil plants of the Carboniferous
period, and occur also in the Devonian. They are preserved as striated,
jointed, cylindrical, or compressed stems, with fluted channels or
furrows at their sides, and sometimes surrounded by a bituminous
coating, the remains of a cortical integument. They were originally
hollow, but the cavity is usually filled up with a substance into which
they themselves have been converted. They were divided into joints or
segments, and when broken across at their articulations they show a
number of striæ, originating in the furrows of the sides, and turning
inwards towards the centre of the stem. It is not known whether this
structure was connected with an imperfect diaphragm stretched across the
hollow of the stem at each joint, or merely represented the ends of
woody plates of which the solid part of the stem is composed. Their
extremities have been discovered to taper gradually to a point, as
represented in _C. cannæformis_ (Fig. 64), or to end abruptly, the
intervals becoming shorter and smaller. The obtuse point is now found to
be the root. Calamites are regarded as Equisetaceous plants; later
botanists consider that they belong to an extinct family of plants.
_Sigillariæ_ are the most abundant of all plants in the coal formation,
and were those principally concerned in the accumulation of the mineral
fuel of the Coal-measures. Not a mine is opened, nor a heap of shale
thrown out, but there occur fragments of its stem, marked externally
with small rounded impressions, and in the centre slight tubercles, with
a quincuncial arrangement. From the tubercles arise long ribbon-shaped
bodies, which have been traced in some instances to the length of twenty
feet.
[Illustration: Fig. 63.--Sphenophyllum restored.]
In the family of the Sigillarias we have already presented the bark of
_S. lævigata_, at page 138; on page 157 we give a drawing of the bark of
_S. reniformis_, one-third the natural size (Fig. 65).
In the family of the Asterophyllites, the leaf of _A. foliosa_ (Fig.
66); and the foliage of _Annularia orifolia_ (Fig. 67) are remarkable.
In addition to these, we present, in Fig. 63, a restoration of one of
these Asterophyllites, the _Sphenophyllum_, after M. Eugene
Deslongchamps. This herbaceous tree, like the Calamites, would present
the appearance of an immense asparagus, twenty-five to thirty feet high.
It is represented here with its branches and _fronds_, which bear some
resemblance to the leaves of the ginkgo. The bud, as represented in the
figure, is terminal, and not axillary, as in some of the Calamites.
[Illustration: Fig. 64.--Calamites cannæformis. One-third natural size.]
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